九州大学 研究者情報
論文一覧
荒谷 邦雄(あらや くにお) データ更新日:2018.07.16

教授 /  比較社会文化研究院 環境変動部門 生物多様性講座


原著論文
1. Makoto Ehara, Kimihiko Hyakumura, Ren'ya Sato, Kiyoshi Kurosawa, Kunio Araya, Heng Sokh, Ryo Kohsaka, Addressing Maladaptive Coping Strategies of Local Communities to Changes in Ecosystem Service Provisions Using the DPSIR Framework, Ecological Economics, 10.1016/j.ecolecon.2018.03.008, 149, 226-238, 2018.07, [URL], The Driver-Pressure-State-Impact-Response (DPSIR) framework has been applied to various environmental problems at multiple spatial and temporal scales and attempts have been made to conceptually improve the framework to encompass various stakeholder perspectives. However, recent literature experiences in the field have challenged the inclusive character of the framework applications. In particular, the framework's inability to incorporate the aggregated informal responses of people affected by changes in ecosystem service provisions has not been fully addressed. This limits the framework's validity in categorizing and disseminating information for addressing particular environmental challenges. Herein, we address this problem by analyzing a case study of deforestation and its impact on non-timber forest product collections by rural residents in Cambodia. We incorporate the concept of maladaptive coping strategies into the DPSIR framework and then further elaborate Ness et al.'s (2010) approach of merging the DPSIR framework with Hägerstrand's (2001) system of nested spatial domains. This conceptualizes the incorporation of the aggregated informal responses into the system, as exemplified in the case study..
2. Tatsuya Mishima, Noriko Wada, Ryûtarô Iwata, Hirosi Anzai, Hosoya Tadatsugu, Kunio Araya, Super-protective child-rearing by japanese bess beetles, cylindrocaulus patalis
Adults provide their larvae with chewed and predigested wood, Insects, 10.3390/insects7020018, 7, 2, 2016.06, [URL], Beetles of the family Passalidae (Coleoptera: Scarabaeoidea) are termed subsocial. The insects inhabit rotten wood as family groups consisting of the parents and their offspring. The Japanese species Cylindrocaulus patalis has the lowest fecundity among passalids because siblicide occurs among the first-instar larvae; accordingly, parental care toward the survived larva is the highest among Passalidae. To clarify the nutritional relationships between the parents and their offspring, we investigated their ability to digest three types of polysaccharides that are components of wood (cellulose and β-1,4-xylan) and fungal cell walls (β-1,3-glucan). Although carboxymethyl-cellulase activity was barely detectable, β-xylosidase, β-glucosidase, β-1,4-xylanase and β-1,3-glucanase activities were clearly detected in both adults and larvae. Because the activities of enzymes that digest β-1,3-glucan were much higher than those for degrading β-1,4-xylan, in both adults and larvae, it is concluded that they are mainly fungivorous. Furthermore, these digestive enzymatic activities in second- and third-instar larvae were much lower than they were in adults. Although all larval instars grew rapidly when fed chewed wood by their parents, larvae ceased growing and died when fed only artificially ground wood meals. We conclude that the larvae are assumed to be provided with chewed predigested wood in which β-1,3-glucan is degraded by parental enzymes..
3. Mishima T. and K. Araya, Analysis of polysaccharide digestive enzymatic systems of the larvae of Dorcus rectus rectus., KOGANE, 18, 94-100, 2016.08.
4. Kon M, Araya K. and Y. Johki, Re-evaluation of the taxonomic status of Leptaulax niae Kuwert (Coleoptera, Passalidae)., KOGANE, 18, 76-78, 2016.08.
5. Mishima, T., Wada N., Iwata R. Anzai H. Hosoya T. and K. Araya, Super-Protective Child-Rearing by Japanese Bess Beetles, Cylindrocaulus patalis: Adults Provide Their Larvae with Chewed and Predigested Wood, Insects, 10.3390/, 7, 2016.03.
6. Dai Suzuki, Mitsuhiro Aizu, Satoshi Yoshizawa, Wataru Tanaka, Kunio Araya, Reptile Diversity in Food Markets in Laos, Current Herpetology, 34, 2, 112-119, 2015.08.
7. Yuta Ito, Ayane Harigai, Moe Nakata, Hosoya Tadatsugu, Kunio Araya, Yuichi Oba, Akinori Ito, Takahiro Ohde, Toshinobu Yaginuma, Teruyuki Niimi, The role of doublesex in the evolution of exaggerated horns in the Japanese rhinoceros beetle, EMBO Reports, 10.1038/embor.2013.50, 14, 6, 561-567, 2013.06, [URL], Male-specific exaggerated horns are an evolutionary novelty and have diverged rapidly via intrasexual selection. Here, we investigated the function of the conserved sex-determination gene doublesex (dsx) in the Japanese rhinoceros beetle (Trypoxylus dichotomus) using RNA interference (RNAi). Our results show that the sex-specific T. dichotomus dsx isoforms have an antagonistic function for head horn formation and only the male isoform has a role for thoracic horn formation. These results indicate that the novel sex-specific regulation of dsx during horn morphogenesis might have been the key evolutionary developmental event at the transition from sexually monomorphic to sexually dimorphic horns..
8. Kiyoto Maekawa, Masahiro Kon, Kunio Araya, New species of the genus Salganea (Blattaria, Blaberidae, Panesthiinae) from Myanmar, with molecular phylogenetic analyses and notes on social structure, Entomological Science, 10.1111/j.1479-8298.2005.00106.x, 8, 1, 121-129, 2005.03, [URL], A new species of the genus Salganea is described from Mount Victoria, Chin State, Myanmar under the name of Salganea matsumotoi sp. nov. Morphological investigation suggests that this species belongs to the S. nigrita species group. The inferred molecular phylogenetic tree indicates that S. matsumotoi is most closely related to the taxa S. incerta, S. taiwanensis, S. gressiti and S. esakii, distributed in Thailand, Taiwan and Japan, all of which belong to the S. nigrita species group. Social structural data suggest that this species is subsocial, and in contrast to other studied species in the genus, potentially has more than one reproductive episode..
9. Kiyoto Maekawa, Masahiro Kon, Tadao Matsumoto, Kunio Araya, Nathan Lo, Phylogenetic analyses of fat body endosymbionts reveal differences in invasion times of blaberid wood-feeding cockroaches (Blaberidae
Panesthiinae) into the Japanese archipelago, Zoological Science, 10.2108/zsj.22.1061, 22, 10, 1061-1067, 2005.10, [URL], Cockroaches have endosymbiotic bacteria in their fat bodies. Recent molecular phylogenetic analyses on both hosts and endosymbionts have revealed that co-evolution has occurred throughout the history of cockroaches and termites. Co-cladogenesis was also shown among closely related taxa (woodroach genus Cryptocercus; Cryptocercidae), and thus endosymbiont data are likely to be useful for biogeographical analyses. To test the possibility of co-cladogenesis among inter- and intraspecific taxa, as well as the utility of endosymbiont data for inferring biogeographical scenarios, we analyzed rRNA genes of endosymbionts of Japanese and Taiwanese Panesthiinae (Salganea and Panesthia; Blaberidae), on which phylogenetic analyses previously had been performed based on the mitochondrial genes. Statistical analyses on the topologies inferred from both endosymbiont and host mitochondria genes showed that co-cladogenesis has occurred. The endosymbiont sequences examined appear to have evolved in a clock-like manner, and their rate of evolution based on the host fossil data showed a major difference in the time of invasion of the two Japanese genera, that is congruent with the recent analyses of their mitochondrial genes..
10. Hosoya Tadatsugu, Kunio Araya, Phylogeny of Japanese stag beetles (Coleoptera
Lucanidae) inferred from 16S mtrRNA gene sequences, with reference to the evolution of sexual dimorphism of mandibles, Zoological Science, 10.2108/zsj.22.1305, 22, 12, 1305-1318, 2005.12, [URL], As a first step in reconstructing the phylogeny of world stag beetles (Coleoptera: Lucanidae), phylogenetic relationships among the major members of Japanese stag beetles were explored by analyzing a sequence of 1030 nucleotides from the mitochondrial 16S ribosomal RNA (16S rRNA) gene. A total of 20 species and three additional subspecies representing 13 genera were examined to provide basic information on the phylogeny of world Lucanidae. The resultant phylogenetic tree indicates that the family Lucanidae is monophyletic, and contains two major lineages: one consists of the genera Platycerus, Aesalus, Ceruchus, and Nicagus, and the other includes Dorcus, Rhaetulus, Prosopocoilus, Aegus, Neolucanus, Prismognathus, Lucanus, Figulus, and Nigidius. Generic members of the latter lineage are further divided into the following four sublineages: i) Figulus and Nigidius; ii) Prismognathus and Lucanus; iii) Aegus and Neolucanus; and iv) Dorcus, Rhaetulus, and Prosopocoilus. These molecular phylogenetic relationships are used as a basis for a preliminary exploration of the evolution of sexual dimorphism in the shape of the mandible. The results of this investigation suggest that strong sexual dimorphism with well-developed mandibles in males evolved independently at least twice, once in the genus Aegus and once in the ancestor of the Lucanus-Prismognathus and Dorcus-Rhaetulus- Prosopocoilus clades. Alternatively, it is possible that sexual dimorphism of mandibles has undergone secondary loss in the genera Figulus and Nigidius..
11. Kiyoto Maekawa, Masahiro Kon, Tadao Matsumoto, Osamu Kitade, Kunio Araya, Phylogeography of the Asian wood-feeding cockroach Salganea raggei roth (Blattaria
Blaberidae) based on the mitochondrial coii gene, Oriental Insects, 41, 317-325, 2007, The mainland Asian species of the wood-feeding cockroach, Salganea raggei Roth, has the widest distribution range in the genus, extending from the eastern Himalayas to Taiwan. The phylogeny of S. raggei was studied in relation to its geographic distribution based on the complete mitochondrial cytochrome oxidase subunit II for 16 representatives collected from variable localities as well as the other Salganea species and outgroup taxa. The resultant phylogenetic tree showed the monophyly of S. raggei and, furthermore, revealed some intraspecific monophyletic groups for this species. It was also revealed that the ancestors of the Nepalese population (the westernmost one) separated from the others, the western Myanmar population (the second westernmost one) followed, and the remaining group split into three monophyletic groups. These results suggest that S. raggei originated around the westernmost part of the present distribution range and, thereafter, spread eastwards yielding derived populations. Preliminary calibration rates based on the COII transversion divergence suggested that the ancestors of S. raggei evolved following the collision between the Indian and Asian plates (∼50 million years ago), and that each monophyletic group was formed from the middle of the Miocene to the Pliocene..
12. Kyoko Ento, Kunio Araya, Shin Ich Kudo, Trophic egg provisioning in a passalid beetle (Coleoptera), European Journal of Entomology, 105, 1, 99-104, 2008, Trophic eggs, which are not viable and eaten by larvae, are produced by the passalid beetle Cylindrocaulus patalis. This is the first record of trophic eggs in subsocial Coleoptera. There are differences in the morphology of trophic and fertile eggs; the former are a paler colour and softer than the latter. The surface of the chorion of trophic eggs is also smoother than that of fertile eggs. The trophic eggs are fed directly by the female parent to 3rd instar larvae following a series of specific behavioural interactions between them, including repeated stridulation by the larva. It is likely that trophic eggs supplement the protein-poor diet of the larvae and contribute to their growth and survival. The production of trophic eggs may be associated with the evolution of an extremely small clutch size in C. patalis..
13. Masaaki Ito, Hisashi Kajimura, Keiko Hamaguchi, Kunio Araya, Ferenc Lakatos, Genetic structure of Japanese populations of an ambrosia beetle, Xylosandrus germanus (Curculionidae
Scolytinae), Entomological Science, 10.1111/j.1479-8298.2008.00280.x, 11, 3, 375-383, 2008.09, [URL], We examined the genetic structures of 13 Japanese populations of an ambrosia beetle, Xylosandrus germanus (Curculionidae: Scolytinae), to understand the effects of geographical barriers on the colonization dynamics of this species. The genetic structure was studied using portions of the mitochondrial cytochrome oxidase I (COI) gene. A phylogenetic analysis revealed three distinct lineages (clades A, B and C) within X. germanus. Clade A contained 21 haplotypes from all 13 populations; whereas clade B contained eight haplotypes from Hokkaido (Sapporo and Furano), Iwate and Nagano populations; and clade C contained only a single a haplotype from the Hokkaido (Furano) population. In the analysis of molecular variance (amova), the greatest amount of genetic variation was detected between populations in Hokkaido and those in Honshu and other southern islands. Between these two groups of populations, all the values of the coefficient of gene differentiation were significantly larger than zero, except for the Hokkaido (Sapporo) versus Nagano comparison. Our results confirm that for X. germanus, gene flow has been interrupted between Hokkaido and Honshu since the last glacial maximum..
14. Kyoko Ento, Kunio Araya, Shin ichi Kudo, Laboratory observation of siblicide with hatching asynchrony in an insect with parental provisioning, Journal of Ethology, 10.1007/s10164-010-0208-2, 28, 2, 405-407, 2010, [URL], We report laboratory observation of siblicide with hatching asynchrony in the passalid beetle Cylindrocaulus patalis. Parents construct a breeding gallery in rotten logs and show complex care of offspring, including provisioning. Clutch sizes are extremely small, and only a single offspring is raised in a colony in each breeding season. We introduced field-collected colonies into an artificial gallery and recorded the behaviour of colony members. Eggs hatched at 2- to 10-day intervals in each colony. Fatal fighting invariably occurred among the larvae soon after hatching of the junior larva, and a single larva, usually the senior one, survived..
15. Dai Suzuki, Kengo Fuse, Mitsuhiro Aizu, Satoshi Yoshizawa, Wataru Tanaka, Kunio Araya, Bounthob Praxaysombath, Reptile diversity in food markets in Laos, Current Herpetology, 10.5358/hsj.34.112, 34, 2, 112-119, 2015, [URL], Wildlife is one of the most important food resources in Laos, and many species are sold at the local markets, but the fauna of the country remains poorly known. As a result of human impacts, including overharvesting for food consumption, many wild animals in Asia face imminent extinction. Our study focused on reptile species that were sold at local markets in Laos. The data were recorded in February, August, and September 2013. We found 16 reptile species in the local market: eight freshwater turtle species, Cuora amboinensis, Cyclemys oldhamii, Malayemys macrocephala, M. subtrijuga, Sacalia quadriocellata, Siebenrockiella crassicollis, Amyda ornata, and Pelodiscus sinensis, one tortoise species, Manouria impressa, four lizard species, Calotes versicolor, Physignathus cocincinus, Varanus nebulosus and V. salvator, and three snake species, Malayopython reticulatus and two species of Ptyas. Most of these reptile species are used by local people as foods. Two species are recorded for the first time from Laos or the surveyed region. Our results suggested that the cryptic diversity of the reptile fauna of Laos still exists and highlighted the importance of field surveys and conservation actions..
16. Kunio Araya, Relationship between the Decay Types of Dead Wood and Occurrence of Lucanid Beetles (Coleoptera
Lucanidae), Applied Entomology and Zoology, 10.1303/aez.28.27, 28, 1, 27-33, 1993, [URL], Relationship between the decay types (white, brown and soft rot) in dead wood and occurrence of lucanid beetles (Coleoptera: Lucanidae) was studied in the field. Of the 292 wood pieces examined, 223 (76.4%) were classified with white rot, 29 (9.9%) with brown rot and 24 (8.2%) with soft rot. The rest were unclassifiable. A total of 595 lucanid beetles (larvae and newly emerged adults) belonging to 8 species (6 genera) were collected from the examined wood. No significant relationship with the decay types was observed in most lucanid species (Prismognathus angularis, Macrodorcas striatipennis, Nipponodorcus montivagus, N. rubrofemoratus and Platycerus delicatulus). However, the occurrence of Ceruchus lignarius and Aesalus asiaticus was significantly concentrated in brown rot, and that of Platycerus acuticollis in soft rot. Moreover, significant positive associations were observed between P. angularis and M. striatipennis and between P. angularis and N. rubrofemoratus, whereas no significant interspecific association was observed for the rest of the species pairs..
17. Kunio Araya, Chemical Analyses of the Dead Wood Eaten by the Larvae of Ceruchus lignarius and Prismognathus angularis (Coleoptera
Lucanidae), Applied Entomology and Zoology, 10.1303/aez.28.353, 28, 3, 353-358, 1993, [URL], The contents of chemical components (alkaline extractives, ethanol/benzene extractives, holocellulose and lignin) were determined for the dead wood in which the larvae of Ceruchus lignarius (brown rot specialist) or Prismognathus angularis (decay type generalist) occurred. The variations of chemical components were small in the samples of C. lignarius and relatively large in those of P. angularis. It was established that C. lignarius occurred exclusively in the brown rotten wood in advanced stage of decay, whereas the occurrence of P. angularis was not as clearly associated with either decay type or its stage..
18. Masafumi Matsui, Hoi Sen Yong, Kunio Araya, Abang Abdul Hamid, Acoustic characteristics and systematic relationships of arboreal Microhylid frogs of the genus Metaphrynella from Malaysia, Journal of Herpetology, 30, 3, 424-427, 1996.12.
19. Ryûtarô Iwata, Fusao Yamada, Hirofumi Katô, Hiroshi Makihara, Kunio Araya, Hisashi Ashida, Masashi Takeda, Nature of galleries, durability of boring scars, and density of Xylotrechus villioni (Villard) larvae (Coleoptera
Cerambycidae), on coniferous tree trunks, Pan-Pacific Entomologist, 73, 4, 213-224, 1997.10, Spatial distributions and shapes of "whirl-like" scars on the trunks, made by gallery formation of mature larvae of Xylotrechus villioni (Villard) (Coleoptera: Cerambycidae), a primary borer of Abies and Picea coniferous trees in Japan, were investigated at an Abies firma Sieb. et Zucc. plantation in Hachiôji, Tokyo Pref., an A. firma natural stand in Miyama, Kyoto Pref. and an A. sachalinensis (Fr. Schm.) Mast. plantation in Imakane, Hokkaidô. Although all the forests investigated showed cumulative "whirl-like" scars on the tree trunks, a low density of existing larvae was inferred from the analyses of the locations and shapes of these scars. Mortality throughout the larval stages, as well as between the final phase of larva and the adult emergence, was suggested. Trunk analysis of a damaged A. firma tree showed that a "whirl-like" scar can remain on the trunk surface for as long as 27 years after the formation of the larval gallery. The most susceptible class of Abies trees had a diameter at the breast height of 35-45cm. "Whirl-like" scars were distributed more densely in the lower part of the trunks..
20. Kunio Araya, Masahiro Tanaka, Luca Bartolozzi, Taxonomic review of the genus Aesalus (Coleoptera
Lucanidae) in the Himalayas, European Journal of Entomology, 95, 3, 407-416, 1998, The genus Aesalus F., 1801 of the family Lucanidae in the Himalayas is reviewed, and a new species, A. saburoi sp. n., is described from Central Nepal. This new species has been confused with A. himalayicus Kurosawa 1985, but differs from the latter mainly by a darker and broader body, and by different shapes of the intercoxal process of the prosternum anbd of male genitalia. The male of A. himalayicus sensu stricto is described for the first time and the male genitalia are illustrated. A key to the members of the genus Aesalus from the Himalayas is provided..
21. Kiyoto Maekawa, Masahiro Kon, Kunio Araya, Tadao Matsumoto, Phylogeny and biogeography of wood-feeding cockroaches, genus Salganea Stål (Blaberidae
Panesthiinae), in Southeast Asia based on mitochondrial DNA sequences, Journal of Molecular Evolution, 10.1007/s002390010252, 53, 6, 651-659, 2001, [URL], Molecular phylogenetic relationships among 25 species of the wood-feeding cockroach belonging to the genus Salganea Stål (Panesthiinae; Blaberidae) in Southeast Asia were analyzed based on the DNA sequence of the complete mitochondrial cytochrome oxidase II (COII) gene. Most basal relationships among species of Salganea are poorly resolved by both neighbor-joining and nonweighted parsimony analyses, suggesting the possibility of a hard polytomy due to a rapid and potentially simultaneous radiation early in the history of the genus. For more apical relationships, however, some interesting phylogenetic relationships were recognized. The monophyly of the two species groups, morio and foveolata, the former of which is distributed mainly in the Sunda lands (containing the Malay Peninsula, Sumatra, Java, and Borneo), whereas the latter is Sulawesi endemic, was strongly supported. Based on the inferred phylogenetic patterns and recent palaeogeographic scenario for Southeast Asia, it is suggested that a radiation of Salganea species occurred in Southeast Asia presumably in the early Tertiary, and several barriers against dispersal and gene flow, such as the formation of straits or high mountains, have arisen from the middle Tertiary..
22. Kyoko Ento, Kunio Araya, Masahiro Kon, The Life History and Colony Composition of Cylindrocaulus patalis (Coleoptera
Passalidae), Sociobiology, 42, 3, 795-806, 2003, The life history and colony composition of Cylindrocaulus patalis (Lewis, 1883) was investigated in Miyazaki Prefecture, Kyushu, southwest Japan, from May 2001 to October 2002. Cylindrocaulus patalis lives in family groups in galleries excavated in rotten wood. Adults were active from April to October and appeared to hibernate from December to March. The breeding season spanned June to August, the hot time of the year. Most observed colonies included a male and female pair year-round. Each pair raised only one offspring per year. Adult off-spring appeared to stay with their parents until the following spring; some stayed until the next breeding season..
23. 荒谷 邦雄, 福岡市早良区の背振山麓で採集されたアトラスオオカブトの生体, Pulex(日本昆虫学会九州支部会報), 94, 682-683, 2015.12.
24. 荒谷 邦雄, 小田切顕一, 田中良尚, 前田芳之, アマミマルバネクワガタの分布に関する新知見, Pulex(日本昆虫学会九州支部会報), 94, 681-682, 2015.12.
25. Yutaka Joki, Masahiro Kon, Kunio Araya, Studies on Leptaulax anibarbis Kuwert (Coleoptera,Passalidae) and its related species, KOGANE (The Japanese Society of Scarabaeoidology), 17, 131-134, 2015.07.
26. Masahiro Kon, Kunio Araya, Yutaka Joki, A new species of the genus Cylindrocaulus (Passalidae, Coleoptera) from Arunachal Pradesh, Northeast India , KOGANE (The Japanese Society of Scarabaeoidology), 17, 127-130, 2015.07.
27. Kunio Araya, Kazuhiro Fujita, A new species of the genus Aegus from Penang Island,West Malaysia., KOGANE (The Japanese Society of Scarabaeoidology), 17, 123-126, 2015.07.
28. 荒谷 邦雄, 北半球の温帯林に隔離分布するクワガタムシ類の日本と北米の関係性, 昆虫と自然, 50, 14, 5-9, 2015.12.
29. 三枝豊平, 荒谷 邦雄, 区系生物地理学から分子系統地理学へ
 ー日本列島の昆虫相ー , 昆虫と自然, 50, 14, 1-4, 2015.12.
30. 荒谷 邦雄, 琉球列島の自然と世界遺産, 昆虫と自然, 50, 5, 24-28, 2015.04.
31. 荒谷 邦雄, 日本の世界遺産の昆虫と自然, 昆虫と自然, 50, 5, 2-4, 2015.04.
32. 荒谷 邦雄, 国内外来種リュウキュウツヤハナムグリを東京都江東区辰巳の森緑道公園で採集, Pulex(日本昆虫学会九州支部会誌), 93, 650-651, 2014.12.
33. 荒谷 邦雄, クレタ島のマツの枯株から発生していたヨーロッパサイカブト., Pulex (日本昆虫学会九州支部会誌), 93, 651-652, 2014.12.
34. K. Araya, A redescription of Hexarthrius sanuchi Fukinuki, 2004 (Coleoptera, Lucanidae) from Cambodia., KOGANE, 16, 2014.07.
35. 荒谷 邦雄, 細谷 忠嗣, 伊豆諸島の魅力を探る~クワガタムシ科を題材に~, 昆虫と自然, 49, 3, 10-17, 2014.03.
36. 荒谷 邦雄, 名古屋議定書が甲虫研究にもたらす影響 〜「名古屋議定書に関する学術関係者意見交換会」に参加して〜 , さやばね(日本甲虫学会), 12, 68-72, 2014.03.
37. 荒谷 邦雄, 大城戸博文, フクロウの巣に発生するコブナシコブスジコガネ(Trox nohirai Nakane)を樹洞中に設置した羽毛トラップで採集., Pulex(日本昆虫学会九州支部会報誌), 92, 624-625, 2014.01.
38. Toyohai Saigusa, Mika Sugimoto, M. Seino, Kunio Araya, On the definition of the psychid genus Bambalina, with notes on Bambalina sp. from Japan (Lepidoptera, Psychidae, Oiketicinae)., Lepidoptera Science, 64, 3, 115-122, 2013.12.
39. Kunio Araya, Tatsuya Mishima, Passalid betles (Coleoptera, Passalidae) collected from Phnom Bokor National Park in southern Cambodia, with a new record of Leptaulax pacholatkoi Iwase from Cambodia., KOGANE, 14, 143-147, 2013.07.
40. 荒谷 邦雄, 伊豆諸島からのカミキリムシ2種の記録, さやばね(日本甲虫学会和文誌), 10, 29-30, 2013.06.
41. Neumann, C., Masahiro Kon, Kunio Araya, Checklist of Passalidae Leach, 1815 (Coleoptera, Scarabaeoidea) of Laos with a key to their identification and a description of Leptaulax pacholatkoi mutoniatus ssp. nov., Entomologica Basiliensia et Collectionis Frey, 34, 207-235, 2013.07.
42. 荒谷 邦雄, ABS問題に楽観視は禁物である ~「 名古屋議定書に関する学術関係者意見交換会」で垣間見えた危機的状況~, 昆蟲(ニューシリーズ), 17, 1, 39-46, 2014.03.
43. Yuta Ito, Ayane Harigai, Moe Nakata, Kunio Araya, Hosoya Tadatsugu, Teruyuki Niimi, The role of doublesex in the evolution of exaggerated horns in the Japanese rhinoceros beetle, EMBO REPORTS, 10.1038/embor.2013.50, 14, 6, 561-567, 2013.04, Male‐specific exaggerated horns are an evolutionary novelty and have diverged rapidly via intrasexual selection. Here, we investigated the function of the conserved sex‐determination gene doublesex (dsx) in the Japanese rhinoceros beetle (Trypoxylus dichotomus) using RNA interference (RNAi). Our results show that the sex‐specific T. dichotomus dsx isoforms have an antagonistic function for head horn formation and only the male isoform has a role for thoracic horn formation. These results indicate that the novel sex‐specific regulation of dsx during horn morphogenesis might have been the key evolutionary developmental event at the transition from sexually monomorphic to sexually dimorphic horns..
44. 荒谷 邦雄, ミクロネシアのコガネムシ上科甲虫相, 昆虫と自然, 47, 9, 5-8, 2012.09.
45. 荒谷 邦雄, 三島達也, ボルネオのクワガタムシ・クロツヤムシ相の概観, 昆虫と自然, 47, 7, 7-10, 2012.07.
46. 荒谷 邦雄, 外来カブトムシ・クワガタムシの生態リスクと管理へのDNAバーコーディングの応用, 昆虫と自然, 47, 1, 16-19, 2012.01.
47. 荒谷 邦雄, 深刻化するペット甲虫の外来種問題, 昆虫と自然, 47, 1, 2-3, 2012.01.
48. Araya K., Notes on Eurytrachelus dorcoides Nagel, 1939 from Sumatra, a junior synonym of Dorcus rama (Boileau, 1897) (Coleoptera, Lucanidae) , KOGANE, 1, 375-383, 2012.08.
49. Araya K. , Redescription of the Male of Aegus hikidai Araya, 1994 Originally Described from Mt. Mulu, Sarawak, Malaysia in Borneo. , Spec. Publ. Jpn.Soc. Scarabaeoidology, 1, 375-383, 2011.10.
50. Araya K. and M. Fijioka, Notes on the Third New Species of the Genus Chewlucanus Ikeda et Katsura, 2000 (Coleoptera, Lucanidae) described from North Sumatra., KOGANE, 12, 2, 375-383, 2011.07.
51. Araya K. and Y. Murai, Description of the second species of the genus Chewlucanus Ikeda et Katsura, 2000 (Coleopreta, Lucanidae) from Sarawak, Malaysia in Borneo., Entomological Revew of Japan, 65, 2, 375-383, 2010.12.
52. Kudo, S., Ento, K. and Araya, K, Siblicide with hatching asynchrony in an insect with parental provisioning., Journal of Ethology, 2010.01.
53. 荒谷邦雄, 伊豆諸島のクワガタムシ相の特徴とその起源、他の分類群との比較, 日本生態学会関東地区会会報, 2009.12.
54. Araya K. and Y. Murai, An Additional Record of Torynognathus chrysomelinus (Coleoptera, Lucanidae) from the Malay Peninsula, with Description of the Female., Elytra , 2009.11.
55. Hosoya, T., Araya,K. and M.Kon. , Phylogenetic relationships of Aulacocyclinae (Coleoptera, Passalidae) inferred from morphological characters. , Elytra, 36, 2008.11.
56. Araya K. and T. Hosoya, A reassessment of Noseolucanus rugosus Araya et Tanaka, 1998 (Coleoptera, Lucanidae), a junior synonym of Lucanus denticulus (Boucher, 1995). , Kogane, (9): 107-110., 2008.07.
57. Ento, K., Araya, K. and Kudo, S. , Trophic egg provisioning in a passalid beetle (Coleoptera). , European. J. Entomol ., 105(1), 2008.01.
58. Kon, M., Johki, Y., Araya, K and Hosoya, T., The hind wings of the genus Cylindrocaulus (Coleoptera, Passalidae, Aulacocyclinae). , Kogane, (8): 45-50., 2007.07.
59. Johki, Y., Araya, K. and Kon, M. , A new species of the genus Leptaulax (Coleoptera, Passalidae) from Sumatra. , Kogane, (8): 51-54., 2007.07.
60. Araya,K., Hosoya, T. and Shima, H. , Notes on the new habitat and parasite of the larvae of coconut rhinoceros beetle, Oryctes rhinoceros (Coleoptera, Scarabaeidae). , Kogane, (8): 39-43., 2007.07.
61. Araya, K. and Fujioka, M. , A new species of the genus Aegus (Coleoptera, Lucanidae) from South Sulawesi, Indonesia. , Kogane, (8): 103-106., 2007.07.
62. Kunio Araya, Notes on a problematical Himalayan lucanid beetle, Dorcus rudis Westeood in Parry (1864), a junior synonym of Dignophorus elegans (Parry, 1862). , Elytra, 35: 85-90., 2007.05.
63. Maekawa, K., Kon, M.,Matsumoto, T., Kitade, O.,& Araya,K., Phylogeography of the Asian wood-feeding cockroach Salganea raggei Roth based on the mitochondrial COII gene. , Oriental Insects , 41:317-325., 2007.01.
64. Yûki Imura and Kunio Araya,, New Records of Ceruchus and Aesalus (Coleoptera, Lucanidae) from China. , Kogane , (7):81-86, 2006.07.
65. Kon, M., K.Araya and M. Mohamed, Two new species of Aceraius (Coleoptera, Passalidae) from Borneo,with a key to Bornean Species of Aceraius. , Elytra , 34: 107-117., 2006.05.
66. Araya, K.,and T.Hosoya, Molecular phylogeny of the Dorcus velutinus species-group (Coleoptera: Lucanidae) inferred from the mitochondrial 16S rRNA gene sequences with the special reference to the taxonomic status of its Japanese and Taiwanese members., Elytra, 33 (2) : 523-530, 2005.11.
67. Maekawa K, Kon M & Araya K., A new species of the genus Salganea (Blattaria, Blaberidae, Panesthiinae) from Myanmar, with molecular phylogenetic analyses and notes on the social structure., Entomological Science, 8: 121-129, 2005.01.
68. Kon, M., K. Mizota & K. Araya, Passalidae (Coleoptera) from the Crocker Range, Sabah, Malaysia., Journal of Tropical Biology and Conservation, 1: 31-46, 2005.01.
69. Kon, M., K. Araya & Y. Johki, A new subspecies of Leptaulax riekoae (Passalidae, Coleoptera) from Myanmar, with a new record of L. riekoae from Laos., Kogane, 6: 47-50, 2005.01.
70. Hosoya, T., and K. Araya, Molecular Phylogeny of Japanese Stag Beetles (Coleoptera: Lucanidae) Inferred from Nucleotide Sequences of the Mitochondrial 16S rRNA Gene with Reference to the Evolution of Sexual Dimorphism of Mandibles., Zool. Sci., (in press), 2005.01.
71. Maekawa K, Kon M, Matsumoto T, Araya K & Lo N., Phylogenetic analyses of fat body endosymbionts reveal differences in invasion times of blaberid wood-feeding cockroaches (Blaberidae: Panesthiinae) into the Japanese archipelago., Zoological Science, 10.2108/zsj.22.1061, 22, 10, 1061-1067, 22 (in press)., 2005.01.
72. Araya, K., & S. Nomura, A New record of Aesalus satoi (Coleoptera, Lucanidae) from Vietnam., Elytra, 32: 21-22, 2004.01.
73. Kon, M.. and K. Araya, Re-evaluation of the status of pelopides mniszechi (Coleoptera, Passalidae), with redescription based on the Holotype., Elytra, 32: 119-123, 2004.01.
74. Kon, M., Kusakabe Y. and K. Araya, A dung ball likely made by a male of Synapsis tridens (Coleoptera, Scarabaeidae)., KOGANE, (5): 13-15, 2004.01.
75. Kon, M., Maekawa, K., Sakai, K., Kusakabe Y. and K. Araya, The cetoniine beetles, Coilodera penicillata and C. miksici (Coleoptera, Scarabaeidae, Cetoniinae), collected from the galleries of wood-feeding cockroaches (Blattaria, Blaberidae, Panesthiinae, Salganea) in Myanmar., KOGANE, (5): 16-20, 2004.01.
76. Araya K., Description of the male genitalia and antennae of Noseolucanus rugosus Araya et Tanaka, 1998 (Coleoptera, Lucanidae), with reference to its systematic position., KOGANE, (5): 41-44, 2004.01.
77. Yutaka Johki, Y., Araya K., and M. Kon, Passalid beetles (Coleoptera, Passalidae) from the Lesser Sunda I. Descriptions of male genitalia of Leptaulax anna, L. sambawae and L. timoriensis and some new distributional records of L. sambawae and L. timoriensis., KOGANE, (5): 45-51, 2004.01.
78. Kon, M., Mizota K. and K. Araya, Passalidae (Coleoptera) recorded from Crocker Range, Sabah, Malaysia., J. Tropical Biology and Conservation, 1:35-50, 2004.01.
79. K. Ayaya and H. Yoshitomi, Discovery of the Lucanid Genus Aesalus (Coleoptera) in the Indochina Region, with Description of a New Species., Spec. Bull. Jpn. Soc. Coleopterol., 6: 189-199, 2003.01.
80. Kon, M., Y. Johki & K. Araya, Two new species of the genus Aceraius (Coleoptera, Passalidae) from Sumatra, with a key to the Sumatran species of Aceraius., Spec. Bull. Jpn. Soc. Coleopterol., 6: 179-187, 2003.01.
81. Johki, Y., K. Araya & M. Kon, A new species of passalid beetle (Coleoptera, Passalidae) closely related to Leptaulax punctatus and L. schoedli from Sumatra., Spec. Bull. Jpn. Soc. Coleopterol., 6: 173-177, 2003.01.
82. Hosoya, T., Araya K. and Y. Shirota, Molecular phylogeny of Japanese stag beetles of the genus Dorcus (Coleoptera, Lucanidae) and its allied genera inferred from mitochondrial COI gene sequences., Elytra, 31(1): 127-142, 2003.01.
83. Kon, M., Y. Johki & K. Araya, A new species of the genus Leptaulax (Coleoptera, Passalidae) from Ph-Pan, Laos., Elytra, 31: 149-154, 2003.01.
84. Kon, M. & K. Araya, On the microhabitat of Ceracupes yui (Coleoptera, Passalidae) from Taiwan., Elytra, 31: 154-156., 2003.01.
85. Johki, Y. & M. Kon, A new bicolored species of Leptaulax (Coleoptera, Passalidae) from Borneo., Elytra, 31: 385-390, 2003.01.
86. Kyoko Ento, Masahiro Kon and Kunio Araya, Life history and colony composition of Cylindrocaulus patalis (Coleoptera, Passalidae)., Sociobiology, 42:795-806, 2003.01.
87. 荒谷 邦雄, ペットとして輸入される外国産コガネムシ上科甲虫の影響, 森林科学, 38: 15-26, 2003.01.
88. Kon, M., K. Araya & Y. Johki, Relationship between microhabitat and relative body thickness in adult beetles of the Oriental passalid genera Aceraius, Macrolinus and Ophrygonius (Coleoptera, Passalidae)., Spec. Bull. Jpn. Coleopterol., (5): 297-304, 2002.01.
89. Johki, Y., K. Araya & M. Kon, A new record of Leptaulax parvus (Coleoptera, Passalidae) from the Malay Peninsula, with a key to the subspecies., Kogane, 3: 35-38., 2002.01.
90. Kon, M., Y. Johki & K. Araya, Notes on Ophrygonius emas (Iwase, 1998), comb. nov. (Coleoptera, Passalidae)., Elytra, 30: 451-453, 2002.01.
91. Hosoya, T., Araya K. and Y. Shirota, Molecular phylogeny of Japanese and Taiwanese stag beetles of the genus Dorcus and its allied genera (Coleoptera, Lucanidae) based on random amplified polymorphic DNA., Giornale Italiano di Entomologia, 10:133-140, 2002.01.
92. 荒谷 邦雄, 腐朽材の特性がクワガタムシ類の資源利用パターンと適応度に与える影響., 日本生態学会誌, 52 (1): 89-98, 2002.01.
93. Araya, K., Kon, M., M. Tanaka, Notes on the genus Eulucanus Didier, 1927, a junior synonym of the genus Prosopocoilus Hope and Westwood, 1845., Kogane, (2):28-32, 2001.01.
94. Araya, K., New Records of Cerchus lignarius monticola NAKANE, 1978 (Coleoptera, Lucanidae) from Fukui Prefecture, Central Japan, with the Reference to Lucanid Fauna of Fukui Prefecture, Spec. Publ. Japan Coleopt. Soc. Osaka, (1): 271-273, 2001.01.
95. Hosoya, T., Honda, M. and Araya, K., Genetic variation of 16S r RNA gene observed in Ceruchus lignarius and Dorcus rectus rectus (Coleoptera, Lucanidae), Entomol. Sci., 4. 335-344, 2001.01.
96. Kon, M., K. Araya & Y. Johki, Redescription of Pelopides monticulosus (Smith) (Coleoptera, Passalidae) based on the type specimen., Kogane, 2: 37-40, 2001.01.
97. Johki, Y., K. Araya & M. Kon, Some new records of five species of Leptaulax (Coleoptera, Passalidae)., Kogane, 2: 41-44, 2001.01.
98. Kon, M., K. Araya & Y. Johki, Notes on the genus Ceracupes Kaup (Passalidae, Coleoptera) from Vietnam, with a new record of C. arrowi Heller., Elytra, 29: 441-445, 2001.01.
99. K. Maekawa, M. Kon, K. Araya and T. Matsumoto, Phylogeny and biogeography of wood-feeding cockroaches genus Salganea Stal (Blaberidae: Panesthiinae) in Southeast Asia based on mitchondrial DNA sequences., J. Mol. Evol., 10.1007/s002390010252, 53, 6, 651-659, 53 :651-659, 2001.01.
100. Kon, M., K. Araya & Y. Johki, Synopsis of Thai Passalidae (Insecta: Coleoptera)., Nat. Hist. J. Chulalongkorn Univ., 1: 61-67., 2001.01.
101. Araya, K., Notes on Lucanus ritae Lacroix, 1984, a junior synonym of Lucanus datunensis Hashimoto, 1984, Kogane, (1): 1-4, 2000.01.
102. Muramoto R. and K. Araya, A new species of the genus Fruhstorferia Kolbe, 1894 (Scarabaeidae, Rutelinae) from northern Thailand, Kogane, (1): 12-16, 2000.01.
103. Ashida, H. K. Kitayama and K. Araya, A new record of Thalassoduvalius masidai masidai S. Ueno (Coleoptera, Trechinae) from Kanmuri-jima Island, Kyoto Prefecture, Central Japan., Elytra, 28 (1): 37, 2000.01.
104. Kon, M., Tanaka M., Araya, K. and Y. Nose, Passalid beetles (Coleoptera, Passalidae) collected from northern Myanmar, with new records of Ceracupes chingkini and Tiberioides borealis., Elytra, 27 (1): 199-205, 1999.01.
105. Kon, M., M. Tanaka & K. Araya, Passalid beetles (Coleoptera, Passalidae) collected from Bhutan, with description of the male genitalia of Tiberioides austeni., Elytra, 27: 471-474, 1999.01.
106. Kon, M. & K. Araya, Description of the male genitalia of Ophrygonius sternibarbis (Coleoptera, Passalidae), with new records from Mt. Trus Madi, Sabah and Mt. Mulu, Sarawak., Elytra, 27: 475-476, 1999.01.
107. Kon, M., T. Ueno & K. Araya, A new record of Cylindrocaulus davidi (Coleoptera, Passalidae) from Gansu, China., Elytra, 27: 476-477, 1999.01.
108. Kunio Araya, Tadatsugu Hosoya and Masahiro Kon, Phylogeny and sexual dimorphism of stag beetles (Coleoptera, Lucanidae)., In : Advances in Ethology 34. Supplements to Ethology (Shakunthala Sridhara ed.). Blackwell Science, 164, 1999.01.
109. Teruo Ochi, Masahiro Kon & Kunio Araya, Notes on the Coprophagous scarab beetles from South-East Asia(I). A New Genus and Species of the Tribe Canthonini from Sumatra., Ent. Rev. Japan, 52 (2): 111-115, 1998.01.
110. Kunio Araya, Masahiro Tanaka and Luca Bartolozzi, Taxonomic review of the genus Aesalus (Coleoptera: Lucanidae) in the Himalayas., European. J. Entomol, 95:407-416, 1998.01.
111. Yutaka Johki, Kunio Araya & Masahiro Kon, Further notes on the microhabitat of Taeniocerus pygmaeus (Coleoptera, Passalidae)., Elytra, 26 (1) : 141-143, 1998.01.
112. Kunio Araya and Masahiro Tanaka, Discovery of the new genus allied to the genus Lucanus (Coleoptera, Lucanidae) from the high mountain in north Myanmar., Elytra, 26 (2) : 333-339, 1998.01.
113. Kunio Araya & Masahiro Kon, Eurytrachellelus coussmenti Maes, 1982, a new synonym of Dorcus kyanrauensis (Miwa), 1934 (Coleoptera, Lucanidae)., Jpn. J. Syst. Ent, 4 (2):297-300, 1998.01.
114. Kon, M., T. Ochi, J. Nabhitabata, K. Araya and M. Matsui, Necrophagous scarab beetles (Coleoptera, Scarabaeidae, Onthophagus) attracted to a diplopod copulating pair (Diplopoda) in Thailand., Elytra, 26(2): 347-349, 1998.01.
115. Kon, M., Panha, S., Araya, K. and M. Matsui, Passalid beetles (Coleoptera, Passalidae) collected from Thailand, with new records of Ophrygonius cantori (Peracheron) and Aceraius laevicollis (Illiger)., Elytra, 26 (2): 350-352, 1998.01.
116. Masanao Honda, Tsutomu Hikida, Kunio Araya, Hidetoshi Ota, Jarujin Nabhitabhata and Yong Hoi-Sen, Cosymbotus craspedotus (Frilly Gekko)and C. platyurus (Flat-tailed Gekko). Gliding Behavior., Herpetol. Review, 28 (1): 42-43, 1997.01.
117. Kunio Araya, Somsak Panha & Masahiro Kon, On a colony of Ceracupes fronticornis (Coleoptera, Passalidae) observed in northern Thailand, with reference to the known microhabitats of the genus Ceracupes., Elytra, 25 (2): 425-428, 1997.01.
118. Masahiro Kon, Masahiro Tanaka & Kunio Araya, Discovery of Cylindrocaulus davidi Boucher et Reyes-Castello (Coleoptera, Passalidae) from the Daling Shan Range, Sichuan, China., Elytra, 25 (2): 429-430, 1997.01.
119. Ryutaro Iwata, Fusao Yamada, Hirofumi Kato, Hiroshi Makihara, Kunio Araya Hisashi Ashida & Masashi Takeda, Nature of Galleries, durability of boring scars, and density of Xylotrechus villioni (Villard) larvae (Coleoptera, Cerambycidae), on coniferous tree trunks., Pan-Pacific Entomol, 73 (4): 213-224, 1997.01.
120. Teruo Ochi & Kunio Araya, Studies on the coprophagous scarab beetles from east Asia. III (Coleoptera, Scarabaeidae)., Gionale Italiano di Entomologica, 8:1-15. (1996), 1997.01.
121. Kunio Araya, Masahiro Kon & Luca Bartolozzi, Morphological Comparisons of the Old World Aesalini (Insecta, Coleoptera, Lucanidae), with special reference to Aesalus imanishii from Taiwan., In The Symposium on the Phylogeny, Biology and Consevation of Fauna and Flora of East Asian Region, 1997 (Kuang Yang Lue and Tien Hsi Chen eds.). Dept. Biol., National Taiwan Univ. and National Science Council, R.O. C, 125-133, 1997.01.
122. Masahiro Kon, Kunio Araya & Yutaka Johki, On the microhabitat of Taeniocerus pygmaeus (Coleoptera, Passalidae)., Elytra, 24 (1) : 47-48, 1996.01.
123. Kunio Araya, Analysis of wood sugars in the dead wood eaten by the larvae of Ceruchus lignarius and Prismognathus angularis (Coleoptera: Lucanidae) by gas-liquid chromatography., Elytra, 24 (2):293-297, 1996.01.
124. Kunio Araya & Abang Abdul Hamid, Discovery of Aesalus yongi (Coleoptera, Lucanidae) from Borneo., Elytra, 24 (2):329-330, 1996.01.
125. Matsui, M., H.-S. Yong, K. Araya and A. A. Hamid, Acoustic characteristics and systematics relationships of arboreal microhylid frogs of the genus Metaphrynella from Malaysia., J. Herpetol, 30 (3): 424-427, 1996.01.
126. Kunio Araya, A redescription of Aesalus himalayicus (Coleoptera, Lucanidae) from Nepal., Spec. Bull. Jpn. Soc. Coleopterol., Tokyo, 365-370, 1995.01.
127. Kunio Araya, Masafumi Matsui, Jarujin Nabhitabhata and Somsak Panha, A New Bristly Aesalus (Coleoptera, Lucanidae) from Peninsular Thailand., G. it. Ent., 7: 73-77 (1994), 1995.01.
128. Kunio Araya, Masahiro Tanaka and Motohiko Tanikado, Discovery of the Lucanid Genus Aesalus (Coleoptera) in the mainland China, with Description of a New Species., Elytra, 23 (2): 93-107, 1995.01.
129. Akira Mori, Kunio Araya and Tsutomu Hikida, Biology of the Poorly Known Bornean Lizard, Apterygodon vittatus, (Squamata, Scincidae)An Arboreal Ant-Eater., Herpetological Natural History, 3 (1): 1-14, 1995.01.
130. Kunio Araya, Toru Kikuta & Tsuyoshi Okuma, The genus Penichrolucanus (Coleoptera, Lucanidae) new to the fauna of Borneo., Elytra, 22 (1): 151-152, 1994.01.
131. Kunio Araya, Oviposition preference of Ceruchus lignarius (Coloptera, Lucaidae) to decay types of dead wood., Japanese Journal of Entomology, 62 (2): 361-362, 1994.01.
132. Kunio Araya, Discovery of a flightless Aegus (Coloptera, Lucaidae) in Borneo., Elytra, 22 (2): 271-280, 1994.01.
133. Masahiro Kon & Kunio Araya, A new record of Opfrygonius singapurae (Coleoptera, Passalidaea) from Thailand., Elytra, 22 (2): 280, 1994.01.
134. Masahiro Kon, Kunio Araya & Yutaka Johki, Aceraius (Coleoptera, Passalidae) new to the fauna of the Korean Peninsula., Elytra, 22 (2): 314, 1994.01.
135. Kunio Araya, Relationship between the decay types of dead wood and occurrence of lucanid beetles (Coleoptera: Lucanidae)., Applied Entomology and Zoology,, 28 (1): 27-33, 1993.01.
136. Kunio Araya, Terra Ochi & Yutaka Johki,, The trogid genus Xizangia Zhang, 1988, a junior synonym of the lucanid genus Penichrolucanus Deyrolle, 1863., Elytra, 21 (1): 87-91, 1993.01.
137. Kunio Araya, Masahiro Kon & Yutaka Johki, A new species of the genus Aesalus (Coleoptera, Lucanidae) from the Malay Peninsula., Elytra, 21 (1): 93-107, 1993.01.
138. Kunio Araya, Notes on the scientific name of some lucanid genera (Coleoptera, Lucanidae)., Elytra, 21 (1): 109-110, 1993.01.
139. Masahiro Kon, Kunio Araya & Yutaka Johki, Passalid beetles (Coleoptera, Passalidae) collected from Sarawak, Borneo. 1. The subfamily Passalinae, with description of a new Aceraius species from Mt. Mulu., Elytra, 21 (1): 115-122, 1993.01.
140. Kunio Araya, Chemical analyses of the dead wood eaten by the larvae of Ceruchus lignarius and Prismognathus angularis (Coleoptera: Lucanidae)., Applied Entomology and Zoology, 28 (3): 353-358, 1993.01.
141. Kunio Araya, Size comparison of Prismognathus angularis (Coleoptera, Lucanidae) collected from brown and white rotten wood., Japanese Journal of Entomology, 61 (3): 562, 1993.01.
142. Kunio Araya, Two new species of tropical Aesalus (Coleoptera, Lucanidae) from the Malay Peninsula., Japanese Journal of Entomology, 61 (4): 697-710, 1993.01.
143. Masahiro Kon, Kunio Araya & Yutaka Johki, A new species of Aceraius (Coleoptera, Lucanidae) from Sabah, Borneo, with redescription of A. moeschleri Kuwert., Japanese Journal of Entomology, 61 (4): 711-717, 1993.01.
144. Kunio Araya, Records of some Lucanidae and Scarabaeidae (Coleoptera) from Minna-jima and Sesoko-jima Islands of Okinawa Group, Ryukyu Archipelago., Elytra, 20 (1): 61-66, 1992.01.
145. Masahiro Kon, Kunio Araya & Yutaka Johki,, A new record of Ophrygonius rajanus (Coleoptera, Passalidae) from the Malay Peninsula., Elytra, 20 (1): 122, 1992.01.
146. Masahiro Kon & Kunio Araya, On the microhabitat of the Bornean passalid beetle, Taeniocerus (Coleoptera, Passalidae)., Elytra, 20 (1): 129-130, 1992.01.
147. Ryutaro Iwata, Kunio Araya & Yutaka Johki, The community of arthropods with spherical postures, including Madrasostes kazumai (Coleoptera: Ceratocanthidae), found from the abandoned part of a nest of Coptotermes formosanus (Isoptera: Rhinotermitidae) in Tokara-Nakanoshima Island, Japan., Sociobiology, 20 (3): 233-244, 1992.01.
148. Masahiro Kon, Kunio Araya & Yutaka Johki, A new species of Aceraius (Coleoptera, Passalidae) from the Malay Peninsula., Elytra, 20 (2): 203-206, 1992.01.
149. Kunio Araya & Teruo Ochi, A new record of Aesalus timidus (Coleoptera, Lucanidae) from the Malay Peninsula., Elytra, 20 (2): 206, 1992.01.
150. Teruo Ochi & Kunio Araya, Studies on the coprophagous scarab beetles from east Asia. II (Coleoptera, Scarabaeidae)., Gionale Italiano di Entomologica, 6: 79-108, 1992.01.
151. Kunio Araya, On the larvae of lucanid beetle, Macrodorcas rectus (Coleoptera, Lucanidae), collected from fallen moso bamboos., Elytra, 19 (2): 271-272, 1991.01.
152. Masahiro Kon & Kunio Araya, On a colony of the Bornean passalid beetle, Leptaulax bicolor (Coleoptera, Passalidae), collected from a fallen bamboo., Japanese Journal of Entomology, 59 (2): 467-468., 1991.01.
153. Kunio Araya, The oviposition mark of the Bornean lucanid beetle, Prosopocoilus occipitalis (Coleoptera, Lucanidae)., Japanese Journal of Entomology, 59 (3): 693-694, 1991.01.

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