溶液内の特に生体分子の分子認識機構、電子移動理論、水溶液の誘電的性質、
溶液構造とゆらぎ、溶液中での熱力学量の評価法、細胞質での混み合い問題、
同符号電荷間実効引力の理論および実験、巨大分子の拡散現象の理論的研究、選択的吸蔵の理論および実験、
キーワード：溶液論、混み合い問題、分子認識、分光学、生体分子、ソフトマター、高分子電解質、選択的吸蔵
2003.03.

主要著書

1.	安池智一, 秋山 良, 改訂版　エントロピーからはじめる熱力学, 放送大学教育振興会（NHK出版）, 2020.03.
2.	安池智一, 秋山 良, エントロピーからはじめる熱力学, 放送大学教育振興会（NHK出版）, 2016.03.

主要原著論文

1.	M. Takeda, K. Maruyama, R. Akiyama and T. Miyata, Integral equation study of effective attraction between like-charged particles mediated by cations: Comparison between IPY2 and HNC closures, EPL (Europhys Letter), doi: 10.1209/0295-5075/ac94f5, 140, 17001-p1-17001-p7, 2022.10, Effective interactions between like-charged particles immersed in an electrolyte solution were calculated using two integral equation theories, hypernetted-chain (HNC)-Ornstein- Zernike (OZ) and ionic Percus-Yevick 2 (IPY2)-OZ. When the HNC-OZ theory was adopted, the electrolyte concentration dependence of the effective interaction showed a reentrant behavior. By contrast, the IPY2-OZ theory did not indicate the behavior. Monte Carlo simulations were performed for one of the model systems, and the results agreed qualitatively with those calculated using the HNC-OZ theory..
2.	Tomoya Iwashita, Masaaki Nagao, Akira Yoshimori, Masahide Terazima, Ryo Akiyama, Usefulness of higher-order system-size correction for macromolecule diffusion coefficients: A molecular dynamics study, Chemical Physics Letters, 10.1016/j.cplett.2022.140096, 807, 140096-1-7, 2022.10, [URL].
3.	Shota Arai, Akira Yoshimori, Yuka Nakamura, Ryo Akiyama, A Microscopic Theory for Preferential Solvation Effects on Viscosity, Journal of Physical Society of Japan, 10.7566/jpsj.91.094602, 91, 9, 094602-1-13, 2022.09, [URL].
4.	Hiraiwa, Tetsuya; Akiyama, Ryo; Inoue, Daisuke; Kabir, Arif Md. Rashedul; Kakugo, Akira, Collision-induced torque mediates the transition of chiral dynamic patterns formed by active particles, PHYSICAL CHEMISTRY CHEMICAL PHYSICS, 10.1039/d2cp03879j, 2022.10, [URL].
5.	Keiju Suda, Ayumi Suematsu, and Ryo Akiyama, Lateral depletion effect on two-dimensional ordering of bacteriorhodopsins in a lipid bilayer: A theoretical study based on a binary hard-disk model, Journal of Chemical Physics, https://doi.org/10.1063/5.0044399, 154, 204904-1-204904-11, 2021.05, [URL].
6.	Suematsu, Ayumi; Akiyama, Ryo, Solvent effect for an effective attraction between like-charged macroions immersed in an electrolyte solution: The intensification mechanism of the effective attraction caused by the translational motion of solvent particles, Journal of Chemical Physics, https://doi.org/10.1063/5.0033874, 154, 3, 034902-1-034902-8, 2021.01, [URL].
7.	Y. Tamura1, A. Yoshimori, A. Suematsu and R. Akiyama, Reentrant crystallization of like-charged colloidal particles in an electrolyte solution: Relationship between the shape of the phase diagram and the effective potential of colloidal particles, EPL (Europhys Letter), 10.1209/0295-5075/129/66001, 129, 66001-p1-66001-p7, 2020.04, The reentrant crystallization of like-charged colloidal particles in electrolyte solutions was studied to clarify the relationship between the shape of the phase diagram and the particle-particle effective potential. Coexisting densities were calculated at various electrolyte concentrations using the thermodynamic perturbation expansion method with effective one-component models. The effective potentials were obtained using an integral equation theory for liquids. Some model effective potentials were examined. The calculated results indicated that the reentrant behaviors of various acidic protein solutions observed in the experiments required not only the nonmonotonic dependence of the short-range attraction on the electrolyte concentration, but also the absence of a long repulsive tail..
8.	Ayano Chiba, Akio Oshima, Ryo Akiyama, Confined Space Enables Spontaneous Liquid Separation by Molecular Size Selective Absorption of Alkanes into a Polyolefin Cast Film, Langmuir, 10.1021/acs.langmuir.9b02509, 35, 52, 17177-17184, 2019.12, [URL], The depletion force has been used to explain phase separation phenomena in colloidal systems. Here, we showed that depletion force can explain not only phase separation of large and small colloidal particles but also preferential absorption of larger molecules from a mixture of large and small molecules in a liquid state. When a polyolefin cast film was immersed in a mixture of long and short normal alkanes, the longer molecules were selectively absorbed into the film. This experimental result was explained from the viewpoint of depletion force. The main finding was the use of confined space to emphasize the separation tendency caused by the force. In general, the increase in entropy may serve as a driving force to mix molecules. However, if sufficiently narrow pores are present, large and small molecules are separated naturally by size as the entropy increases. This finding will lead to size exclusion chromatography of low-mass molecules, similar to gel permeation chromatography of macromolecules. In order to demonstrate the effect of depletion force, we selected and experimented with a system based on a polyolefin isotactic poly(4-methyl-1-pentene) (P4MP1) film and a normal alkane mixture and realized high molecular selectivity. The P4MP1 film we used can be prepared simply by evaporating the solvent from the solution and casting the film. On the basis of the Asakura-Oosawa theory, we concluded that spontaneous and high molecular selectivity is attributed to the depletion force provided by the small sub-nanopores with uniform size in the film..
9.	Ken Tokunaga, Ryo Akiyama, Molecular dynamics study of a solvation motor in a Lennard-Jones solvent, Physical Review E, 10.1103/PhysRevE.100.062608, 100, 6, 062608-1-062608-8, 2019.12, [URL], The motions of a solvation motor in a Lennard-Jones solvent were calculated by using molecular dynamics simulation. The results were analyzed considering the large spatial scale effects caused by the motion of the solvation motor. A reaction site was located on the surface of the solvation motor and the attraction between the reaction site and the solvent molecules was varied for 100 fs. The motion of the motor was driven by solvation changes near the reaction site on the motor. Two finite-size effects were observed in the motion. One was the hydrodynamic effect and the other was the increase in solvent viscosity caused by heat generation. The latter affected not only the displacement of the motor caused by the reaction but also the wave propagation phenomena. Both effects reduced the motor displacement. Heat generation affects the displacement, in particular for small systems. By contrast, the hydrodynamic effect remained even for large systems. An extrapolation method was proposed for the displacement..
10.	Yuka Nakamura, Shota Arai, Masahiro Kinoshita, Akira Yoshimori, Ryo Akiyama, Reduced density profile of small particles near a large particle Results of an integral equation theory with an accurate bridge function and a Monte Carlo simulation, Journal of Chemical Physics, 10.1063/1.5100040, 151, 4, 044506-1-044506-10, 2019.07, [URL], Solute-solvent reduced density profiles of hard-sphere fluids were calculated by using several integral equation theories for liquids. The traditional closures, Percus-Yevick (PY) and the hypernetted-chain (HNC) closures, as well as the theories with bridge functions, Verlet, Duh-Henderson, and Kinoshita (named MHNC), were used for the calculation. In this paper, a one-solute hard-sphere was immersed in a one-component hard-sphere solvent and various size ratios were examined. The profiles between the solute and solvent particles were compared with those calculated by Monte Carlo simulations. The profiles given by the integral equations with the bridge functions were much more accurate than those calculated by conventional integral equation theories, such as the Ornstein-Zernike (OZ) equation with the PY closure. The accuracy of the MHNC-OZ theory was maintained even when the particle size ratio of solute to solvent was 50. For example, the contact values were 5.7 (Monte Carlo), 5.6 (MHNC), 7.8 (HNC), and 4.5 (PY), and the first minimum values were 0.48 (Monte Carlo), 0.46 (MHNC), 0.54 (HNC), and 0.40 (PY) when the packing fraction of the hard-sphere solvent was 0.38 and the size ratio was 50. The asymptotic decay and the oscillation period for MHNC-OZ were also very accurate, although those given by the HNC-OZ theory were somewhat faster than those obtained by Monte Carlo simulations..
11.	Ayumi Suematsu, Takuto Sawayama, and Ryo Akiyama, Effective potential between negatively charged patches on acidic proteins immersed in various electrolyte solutions, The Journal of Chemical Physics, doi: 10.1063/1.5038912, 149, 074105-1-8, 2018.08, Effective interactions between O-sized anions in various electrolyte solutions were calculated by using the integral equation theory with some simple models. The results indicated that only multivalent cations mediated a strong effective attraction between O-sized anions at a certain concentration. The effective interaction turned from repulsive to attractive as the electrolyte concentration increased, and the effective attraction decreased when more electrolyte was added. Moreover, the effective interactions between O-sized anions in the electrolyte solution did not present a long repulsive tail, although the effective attraction caused by the divalent cations appeared. By contrast, the effective attraction mediated by monovalent cations and the reentrant behavior did not appear and the effective interaction was basically repulsive. These behaviors agree with the experimental results for reentrant condensation of acidic proteins in various electrolyte solutions. The calculated results suggest that the dissociated carboxylic acidic groups on the proteins form attractive patches between proteins under certain concentration conditions..
12.	久保田 陽二, Akira Yoshimori, Nobuyuki Matubayasi, Makoto Suzuki, Ryo Akiyama, Molecular dynamics study of fast dielectric relaxation of water around a molecular-sized ion, JOURNAL OF CHEMICAL PHYSICS, 10.1063/1.4769972, 137, 22, 224502-1-224502-4, 2012.12.
13.	Ryo Akiyama, and Ryo Sakata , An Integral Equation Study of Reentrant Behavior in Attractive Interactions between Like-Charged Macroions Immersed in an Electrolyte Solution, J. Phys. Soc. Jpn., 10.1143/JPSJ.80.123602, 80, 123602, 2011.12.
14.	Yoji Kubota and Ryo Akiyama, Fine Structure of the Dielectric Response to a Molecular-Sized Ion in Water, J. Phys. Chem. Lett., 10.1021/jz200571f, 2, 13, 1588–1591, 2011.06, [URL].
15.	Ryo Akiyama, Yasuhito Karino, Hokuto Obama, Ayako Yoshifuku, Adsorption of xenon on a protein arising from the translational motion of solvent molecules, Phys. Chem. Chem. Phys. , 10.1039/b921314g, 12, 3096-3101, 12, 3096-3101 (Communication), 2010.03, [URL].
16.	Ryo Akiyama, Naohiko Fujino, Kouhei Kaneda, and Masahiro Kinoshita, Interaction between like-charged colloidal particles in aqueous electrolyte solution: Attractive component arising from solvent granularity, Cond. Matt. Phys., 10, 587-596 , 2007.12, [URL].
17.	Masahiro Kinoshita, Yuichi Harano, and Ryo Akiyama, Changes in thermodynamic quantities upon contact of two solutes in solvent under isochoric and isobaric conditions, J. Chem. Phys., 125, 244504-1-7 , 2006.12.
18.	Ryo Akiyama, Yasuhito Karino, Yasuhiro Hagiwara, Masahiro Kinoshita, Remarkable Solvent Effects on Depletion Interaction in Crowding Media: Analyses by the Integral Equation Theories, J. Phys. Soc. Jpn., 75, 064804-1-7 , 2006.05.
19.	Merchant, K.A.; Noid, W.G.; Akiyama, R; Finkelstein, I.; Goun, A.; McClain, B.L.; Loring, R.F.; Fayer,M.D., Myoglobin-CO Substate Structures and Dynamics: Spectrally Resolved Stimulated Vibrational Echoes and Molecular Dynamics Simulations, J. Am. Chem. Soc., (2003) Vol.125, p.13804., 2003.11.
20.	Ryo Akiyama and Roger F. Loring, Quantum Solvent and Solute Effects in the Infrared Vibrational echo, J. Phys. Chem. A, (2003) Vol.107, p.8024, 2003.10.
21.	Ryo Akiyama and Roger F. Loring, Vibrational Echoes for Classical and Quantum Solutes,, J. Chem. Phys., Vol. 116, p.4655(2002)., 2002.01.
22.	Ryo Akiyama, Masahiro Kinoshita, and Fumio Hirata:, Free energy profiles of electron transfer at water-electrode interface studied by the reference interaction site model theory, Chem. Phys. Lett., Vol.305, p.251(1999)., 1999.01.
23.	Ryo Akiyama and Fumio Hirata, Theoretical Study for Water Structure at Highly Ordered Surface: Effect of Surface Structure, J. Chem. Phys., Vol.108, p.4904 (1998)., 1998.01.
24.	Ryo Akiyama, Akira Yoshimori, Toshiaki Kakitani, Yasushi Imamoto, Yoshinori Shichida and Yasuyo Hatano, Analysis of Excited State Dynamics of 13-trans-Locked-Bacteriorhodopsin, J. Phys. Chem. A, Vol. 101, p.412 (1997)., 1997.01.
25.	Ryo Akiyama, Toshiaki Kakitani, Yasushi Imamoto, Yoshinori Shichida and Yasuyo Hatano, Bacteriorhodopsin Analyzed by the Fourier Transform of Optical Absorption Spectra,, J. Phys. Chem., Vol. 99, 7147(1995)., 1995.01.

主要総説, 論評, 解説, 書評, 報告書等

1.	秋山 良, エントロピー駆動実効引力相互作用に関する研究, 溶液化学研究会誌, Vol.1, 8-16(2022), 2022.04.
2.	千葉文野、秋山 良, 高分子P4MP1膜への低分子混合溶液からの選択的分子吸蔵 Selective Molecular Absorption from Low Molecular Weight Mixed Solution to a Polymer P4MP1 Film, 高圧力の科学と技術, https://doi.org/10.4131/jshpreview.31.82, Vol.31, No.2, 82-89(2021), 2021.10.
3.	秋山 良, 末松 安由美, 液体の積分方程式理論を用いた電解質溶液内の同符号電荷間実効引力相互作用の研究, 分子シミュレーション学会誌"アンサンブル", Vol.23, No.2, p.p. 103 - 112, 2021.04.
4.	Hisashi Okumura, Masahiro Higashi, Yuichiro Yoshida, Hirofumi Sato, Ryo Akiyama, Theoretical approaches for dynamical ordering of biomolecular systems, Elsevier, 10.1016/j.bbagen.2017.10.001, BBA (Biochimica et Biophysica Acta) - General Subjects 1862 (2018) 212–228 (査読あり), 2018.02, [URL], Background Living systems are characterized by the dynamic assembly and disassembly of biomolecules. The dynamical ordering mechanism of these biomolecules has been investigated both experimentally and theoretically. The main theoretical approaches include quantum mechanical (QM) calculation, all-atom (AA) modeling, and coarse-grained (CG) modeling. The selected approach depends on the size of the target system (which differs among electrons, atoms, molecules, and molecular assemblies). These hierarchal approaches can be combined with molecular dynamics (MD) simulation and/or integral equation theories for liquids, which cover all size hierarchies. Scope of review We review the framework of quantum mechanical/molecular mechanical (QM/MM) calculations, AA MD simulations, CG modeling, and integral equation theories. Applications of these methods to the dynamical ordering of biomolecular systems are also exemplified. Major conclusions The QM/MM calculation enables the study of chemical reactions. The AA MD simulation, which omits the QM calculation, can follow longer time-scale phenomena. By reducing the number of degrees of freedom and the computational cost, CG modeling can follow much longer time-scale phenomena than AA modeling. Integral equation theories for liquids elucidate the liquid structure, for example, whether the liquid follows a radial distribution function. General significance These theoretical approaches can analyze the dynamic behaviors of biomolecular systems. They also provide useful tools for exploring the dynamic ordering systems of biomolecules, such as self-assembly. This article is part of a Special Issue entitled “Biophysical Exploration of Dynamical Ordering of Biomolecular Systems” edited by Dr. Koichi Kato..
5.	久保田 陽二, 秋山 良, 分子スケールの外場で測る誘電率 -溶質分子周囲の水が作り出すポテンシャル揺らぎ-, 分子シミュレーション研究会会誌"アンサンブル", Vol.15, No.1, p.p. 33 - 41, 2013.01.
6.	秋山良, Asakura-Oosawaモデルからはじめる定積過程と定圧過程の違い, 分子シミュレーション研究会会誌"アンサンブル", Vol.13, No.2, p.p. 57-65, 2011.04.
7.	秋山良, 青少年のためのAsakura-Oosawa理論入門, 生物物理, 51(1), p.p.036-040, (2011), 2011.01.
8.	秋山良、狩野康人、木下正弘, 溶媒分子の並進運動由来の巨大分子間引力相互作用と熱測定, 熱測定, 33(3), p.p.104-113, (2006), 2006.06.

主要学会発表等

1.	Ryo Akiyama*, Theoretical Study of Effective Attraction between Like-Charged Particles and the Reentrant Condensation Behavior, The 4th International Conference on Material Research and Innovation (ICMARI 2022), 2022.12.
2.	秋山良, エントロピー駆動の実効相互作用：膜と分離, 電気化学会第８９回大会, 2022.03.
3.	秋山良, Studies on Effective Attractive Interactions Driven by Entropy, エントロピー駆動実効引力相互作用に関する研究, 第４３回　溶液化学シンポジウム, 2021.10.
4.	Tomoya Iwashita, Msaaki Nagao, Akira Yoshimori, Masahide Terazima and Ryo Akiyama*, Diffusion coefficient of one macromolecule in an aqueous solution: the system size　dependence of the viscosity in the estimation method using MD simulations, The 3rd International Conference on Material Research and Innovation (ICMARI 2021), 2021.12.
5.	秋山良, Asakura-Oosawa 理論とその広がり, Variations on a Theory by Asakura and Oosawa, 日本生物物理学会　第５８回　年会, 2020.09.
6.	Ryo Akiyama, Statistical Mechanics Study of Separation of Inert Solvent Mixtures by a Porous Material, The 2nd International Conference on Material Research and Innovation (ICMARI 2019), 2019.12.
7.	Ryo Akiyama, Asakura-Oosawa theory: On the Origin of Excluded Volume Effects in a Crowding Media and the Progress, the 5th International Conference on Molecular Simulation (ICMS2019), 2019.11.
8.	Ayano Chiba, Akio Oshima, Kenzaburo Okubo, and Ryo Akiyama, Separation of inert solvent mixtures accentuated by confined spaces, 6th Japan-Korea International Symposium on Material Science and Technology 2019 (JKMST2019), 2019.08.
9.	Ryo Akiyama, Effective attraction between negatively charged sites on proteins and ordering of proteins in an electrolyte solution, The 4th International Symposium “Dynamical Ordering of Biomolecular Systems for creation of integrated functions”(新学術領域研究『動的秩序と機能』第4回国際シンポジウム), 2015.11.
10.	Ryo Akiyama, Spatiotemporal dynamic ordering regulated by ATP hydrolysis and effective attraction between negatively charged sites in a biofluid, The 53rd Annual Meeting of the Biophysical Society of Japan, Formation of spatiotemporal dynamic ordering mediated by ATP hydrolysis (新学術 領域研究『動的秩序と機能』共催シンポジウム), 2015.09.
11.	秋山 良, 静電相互作用による巨大分子構造形成とイオン交換による制御, 日本化学会　第95回春季年会, 2015.03.
12.	秋山 良, Association of Actin Monomers and Effective Attraction between Like-Charged Colloidal Particles Mediated by Multivalent Cations, 第52回日本生物物理学会年会, 2014.09.
13.	Ryo Akiyama, Shingo Fujihara, Takuto Sawayama, Akira Yoshimori, Interaction between like-charged particles and attractive patches on like-charged proteins, International Meeting on Application of Statistical Mechanics of Molecular Liquid on Soft Matter, 2014.09.
14.	秋山 良, Hydration dynamics and dielectric relaxation of water around an ion, Symposium on Hydrationand ATP Energy 2013, 2013.03.
15.	久保田 陽二, Akira Yoshimori, Nobuyuki Matubayasi, Makoto Suzuki, 秋山 良, Slow Rotational and Fast Dielectric Relaxation of Water around an Ion: A Molecular Dynamics Study, 4th France- Japan Joint Seminar, 2013.01.
16.	秋山 良, 分子動力学シミュレーションで“観測”されたハイパーモバイル水, 合同公開シンポジウム ゆらぎと水 - 生命のエネルギーと機能 の分子機構を探る, 2012.09.
17.	秋山 良, 朝倉―大沢理論から見渡す生体分子間の実効相互作用, 2012年 生物物理若手の会夏の学校, 2012.09.
18.	秋山 良, 同符号のマクロイオン間の強い引力とリエントラントな電解質濃度依存性, 低次元系光機能材料研究会」第1回サマーセミナー(兼 西日本 ナノシート研究会第1回シンポジウム), 2012.07.
19.	久保田 陽二, Akira Yoshimori, Nobuyuki Matubayasi, Makoto Suzuki, 秋山 良, Fast Dielectric Relaxation of Water around an Ion, 6th Mini- Symposiumon Liquids, 2012.06.
20.	秋山良　, 同符号マクロイオン間の実効引力とSupraparticle Chemistry, 日本化学会第９２回春季年会（「高次分子システムのための分子科学：実験と理論の挑戦」(先端ウォッチング講演会)）, 2012.03.
21.	Ryo Akiyama, Reentrant Behavior in Attractive Interaction between Like-Charged Colloidal Particles Immersed in Electrolyte Solution: a Theoretical Study and a Simple Picture, Seminar uber BioNano-Physik / Medizinische Physik at University of Tuebingen, 2011.11.
22.	秋山良　, 水中の溶質が感じる静電ポテンシャルのゆらぎ, 第１５回分子シミュレーション夏の学校, 2011.09.
23.	秋山良　, 単純なモデルで考える定積過程と定圧過程：分子シミュレーションの課題と実験の課題, 第１５回分子シミュレーション夏の学校, 2011.09.
24.	秋山良　, 同符号マクロイオン間の引力相互作用とリエントラントな挙動：生物化学的なコンテクストで, 愛媛大学理学部物理教室談話会, 2011.08.
25.	Ryo Akiyama, Attractive interaction between like-charged macromolecules in an electrolyte solution and motion of amoeba, Department Seminar (Virginia Commonwealth University) , 2011.03.
26.	秋山良, 電子雲が作る共有結合とイオン雲が作る”共有結合”, 九重分光学関連夏季セミナー, 2010.07.
27.	秋山良, 対イオンがつくる同符号マクロイオン間の“共有結合”とアクチン分子間相互作用： HNC-OZ方程式の解, 九州工業大学　情報工学部 生命情報工学科　学科講演会, 2009.12.
28.	秋山良, van der Waals描像と溶液内での生体分子間相互作用, 新潟大学　理学部 自然環境科学科　コロキウム, 2009.06.
29.	秋山良, タンパク質分子間相互作用における水の並進運動効果と熱力学量, 生物物理学会第４６回年会, 2008.12, [URL].
30.	秋山良, 溶媒分子の粒子性と巨大分子間実効相互作用, 第２２回分子シミュレーション討論会, 2008.11, [URL].
31.	R. Akiyama, Effective interaction between like-charged colloidal particles in electrolyte solution: Attractive component arising from solvent granularity, International Symposium on Structure and Reaction Dynamics of Ionic Liquids in Kanazawa, 2008.09, [URL].
32.	秋山良, 溶媒分子の”枯渇”と生体分子間相互作用－水のあたり前な溶媒効果と想定外の結果－, 第3回ＬＳＷシンポジウム　ソフト＆ウェットマターにおける水の役割 , 2008.01, [URL].
33.	秋山良, 鷹揚生物物理化学, 分子科学若手の会　夏の学校２００７, 2007.08, [URL].
34.	R. Akiyama , Crowding Problems in Cytoplasm and Interaction Arising from Translational Motion of Solvent Molecules, International mini-symposium on liquids, 2007.05, [URL].
35.	秋山　良, 細胞質での混み合い問題と溶媒分子の並進運動, 日本物理学会秋季大会, 2006.09.
36.	秋山　良, ◯、o、。からはじめる物理化学:細胞質での生体分子の混み合い問題と溶媒効果, 物理化学セミナー, 2006.09.
37.	秋山良, 赤外振動エコー実験と分子動力学実験に基づく一酸化炭素結合ミオグロビンの A Substatesの構造とダイナミクス, 生理学研究所　生体ダイナミクス研究会, 2003.03.

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学会大会・会議・シンポジウム等における役割

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学術論文等の審査

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