KONDO HISAO
Organization
Faculty of Medical Sciences Department of Molecular Biology Professor
School of Medicine Department of Biomedical Science（Concurrent）
Graduate School of Systems Life Sciences Department of Systems Life Sciences（Concurrent）
Title
Professor
Contact information
Tel
0926426175
Profile
I have been investigating the biogenesis of organelles, e.g., the Golgi apparatus and ER. Especially, we focus on the clarification of the p97ATPase-mediated membrane fusion.
Homepage
External link
Research Areas
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Life Science / Cell biology
Degree
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Ph.D in Medicine
Research History
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2004年-2006年 三菱化学生命科学研究所
2004年-2006年 三菱化学生命科学研究所
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1988年-1997年 京都大学 1999年-2004年 University of Cambridge
Education
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Kyoto University 医学部 医学科
Research Interests・Research Keywords
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Research theme： Molecular mechanism of the Golgi biogenesis.
Keyword： p97ATPase, membrane fusion, Golgi
Research period： 1995.4
Awards
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病態代謝研究会理事長賞
2011.6 病態代謝研究会
Papers
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p97 and p47 function in membrane tethering in cooperation with FTCD during mitotic Golgi reassembly. Reviewed International journal
Yayoi Kaneko, Kyohei Shimoda, Rafael Ayala, Yukina Goto, Silvia Panico, Xiaodong Zhang, Hisao Kondo
EMBO J 40 ( 9 ) e105853 2021.5
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VCIP135 deubiquitinase and its binding protein, WAC, in p97ATPase-mediated membrane fusion. Reviewed International journal
Totsukawa G, Kaneko Y, Uchiyama K, Toh H, Tamura K, Kondo H
EMBO J 30 ( 17 ) 2011.9
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Analysis of nucleotide binding to p97 reveals the properties of a tandem AAA hexameric ATPase. Reviewed International journal
Briggs LC, Baldwin GS, Miyata N, Kondo H, Zhang X, and Freemont PS.
J Biol Chem 2008.4
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p37 is a p97 adaptor required for Golgi and ER biogenesis in interphase and at the end of mitosis. Reviewed International journal
Uchiyama K, Totsukawa G, Puhka M, Kaneko Y, Jokitalo E, Dreveny I, Beuron F, Zhang X, Freemont P, and Kondo H.
Developmental Cell 2007.1
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Conformational changes in the AAA ATPase p97-p47 adaptor complex. Reviewed International journal
Beuron F, Dreveny I, Yuan X, Pye VE, Mckeown C, Briggs LC, Cliff MJ, Kaneko Y, Wallis R, Isaacson RL, Ladbury JE, Matthews SJ, Kondo H, Zhang X, Freemont PS.
EMBO J 2006.1
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Structural basis of the interaction between the AAA ATPase p97/VCP and its adaptor protein p47. Reviewed International journal
Dreveny I, Kondo H, Uchiyama K, Shaw A, Zhang X and Freemont PS.
EMBO J 2004.1
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The localization and phosphorylation of p47 are important for Golgi disassembly-assembly during the cell cycle. Reviewed International journal
. Uchiyama K, Jokitalo E, Lindman M, Jackman M, Kano F, Murata M, Zhang X and Kondo H.
J Cell Biology 2003.1
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VCIP135, a novel essential factor for p97/p47-mediated membrane fusion, is required for Golgi and ER assembly in vivo. Reviewed International journal
Uchiyama K, Jokitalo E, Kano F, Murata M, Zhang X, Canas B, Newman R, Rabouille C, Pappin D, Freemont P and Kondo H.
J Cell Biology 2002.1
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Structure of the AAA ATPase p97. Reviewed International journal
Zhang X, Shaw A, Bates PA, Newman RH, Gowen B, Orlova E, Gorman MA, Kondo H, Dokurno P, Lally J, Leonard G, Meyer H, van Heel M and Freemont PS.
Molecular Cell 2000.1
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Syntaxin 5 is a common component of the NSF- and p97-mediated reassembly pathways of Golgi cisternae from mitotic Golgi fragments in vitro. Reviewed International journal
. Kondo H#, Rabouille C# (# Joint first authorship), Newman R, Hui N, Freemont P and Warren G.
Cell 1998.1
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p47 is a cofactor for p97-mediated membrane fusion. Reviewed International journal
Kondo H, Rabouille C, Newman R, Levine TP, Pappin D, Freemont P and Warren G.
Nature 1997.1
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VCIP135 associates with both the N- and C-terminal regions of p97 ATPase. Reviewed International journal
Nakayama S, Kondo H.
J Biol Chem 300 ( 1 ) 105540 2024.1
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VCIP135 associates with both the N- and C-terminal regions of p97 ATPase
Nakayama, S; Kondo, H
JOURNAL OF BIOLOGICAL CHEMISTRY 300 ( 1 ) 105540 2024.1 （ ISSN:00219258 eISSN:1083-351X ）
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Mitotic phosphorylation of VCIP135 blocks p97ATPase-mediated Golgi membrane fusion. Reviewed International journal
Go Totsukawa, Ayaka Mtsuo, Ayano Kubota, Hisao Kondo
Biochem Biophys Res Commun 433 ( 2 ) 237 - 242 2013.4
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Isolation of a point-mutated p47 lacking binding affinity to p97ATPase. Reviewed International journal
Kaneko Y, Tamura K, Totsukawa G, Kondo H
FEBS Lett 584 2010.11
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Phosphorylation of p37 is important for Golgi disassembly at mitosis. Reviewed International journal
Kaneko Y, Tamura K, Totsukawa G, Kondo H
Biochem Biophys Res Commun 402 2010.11
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The maintenance of the endoplasmic reticulum network is regulated by p47, a cofactor of p97, through phosphorylation by cdc2 kinase. Reviewed International journal
Kano F, Kondo H, Yamamoto A, Tanaka AR, Hosokawa N, Nagata K and Murata M.
Genes Cells 2005.1
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NSF/SNAPs and p97/p47/VCIP135 are sequentially required for cell cycle-dependent reformation of the ER network. Reviewed International journal
Kano F, Kondo H, Yamamoto A, Kaneko Y., Uchiyama K., Hosokawa N, Nagata K and Murata M.
Genes Cells 2005.1
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Cdc2 kinase-dependent disassembly of endoplasmic reticulum (ER) exit sites inhibits ER-to-Golgi vesicular transport during mitosis. Reviewed International journal
Kano F, Tanaka AR, Yamauchi S, Kondo H and Murata M.
Mol Biol Cell 2004.1
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Structure, dynamics and interactions of p47, a major adaptor of the AAA ATPase, p97. Reviewed International journal
Yuan X, Simpson P, McKeown C, Kondo H, Uchiyama K, Wallis R, Dreveny I, Keetch C, Zhang X, Robinson C, Freemont P and Matthews S.
EMBO J 2004.1
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Complete backbone resonance assignments of p47: the 41kDa adaptor protein of the AAA ATPase p97. Reviewed International journal
Yuan X, Simpson P, Kondo H, McKeown C, Dreveny I, Zhang X, Freemont PS and Matthews S.
J Biomol NMR 2004.1
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Progressive myopathy with circulating autoantibody against giantin in the Golgi apparatus. Reviewed International journal
Sahashi K, Ibi T, Ohno K, Nakao N and Kondo H.
Neurology 2004.1
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Motions and negative cooperativity between p97 domains revealed by cryo-electron microscopy and quantised elastic deformational model. Reviewed International journal
Beuron F, Flynn TC, Ma J, Kondo H, Zhang X and Freemont PS.
J Mol Biol 2003.1
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The crystal structure of murine p97/VCP at 3.6A. Reviewed International journal
Huyton T, Pye VE, Briggs LC, Flynn TC, Beuron F, Kondo H, Ma J, Zhang X and Freemont PS.
J Struct Biol 2003.1
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Involvement of the ryanodine-sensitive Ca2+ store in GLP-1-induced Ca2+ oscillations in insulin-secreting HIT cells. Reviewed International journal
Sasaki S, Nakagaki I, Kondo H and Hori S.
Pflugers Arch 2002.1
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Solution structure and interaction surface of the C-terminal domain from p47: a major p97-cofactor involved in SNARE disassembly. Reviewed International journal
Yuan X, Shaw A, Zhang X, Kondo H, Lally J, Freemont PS and Matthews S.
J Mol Biol 2001.1
Books
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p97ATPaseによるゴルジ体・小胞体の形成維持
近藤 久雄（Role：Joint author）
秀潤社 2011.11
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生体の科学
近藤久雄、十津川剛（Role：Joint author）
医学書院 2008.5
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細胞内小器官形成維持のための新たな分子機構
近藤 久雄（Role：Joint author）
羊土社 2007.4
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Oxidative stress in muscular atrophy In: Handbook of Oxidants and Antioxidants in Exercise
Kondo H.（Role：Joint author）
Elsevier Science 2000.1
Presentations
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p55, a novel p47-binding protein, is required for p97/p47-mediated Golgi reassembly at mitosis.
近藤 久雄, 金子弥生
日本分子生物学会 2016.12
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p55, a novel p47-binding protein, is required for p97/p47-mediated Golgi biogenesis.
近藤 久雄, 金子弥生
日本分子生物学会 2013.12
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p97-ATPase mediated membrane fusion Invited
近藤 久雄, 金子 弥生
日本分子生物学会 2013.12
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p97-mediated biogenesis of the Golgi and ER Invited International conference
近藤久雄
The 3rd Internal Symposium Protein Community 2010.9
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細胞内膜融合機構p97/p47経路の細胞周期調節 Invited
近藤久雄
第54回日本細胞生物学会 2001.5
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膜融合機構p97/p47経路の新規因子について Invited
近藤久雄
第24回日本分子生物学会 2001.12
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p97-mediated membrane fusion Invited
第27回日本分子生物学会 2004.11
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p97ATPase-mediated membrane fusion Invited
近藤久雄、十津川剛
第28回日本分子生物学会 2005.12
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p97-mediated biogenesis of the Golgi and ER. Invited International conference
Kondo H.
The 20th IUBMB in Kyoto 2006.6
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ゴルジ体・小胞体形成に関する最近の知見〜特にp97ATPaseによる膜融合の観点から Invited International conference
近藤久雄
第71回日本生化学会中部支部会 2007.5
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p97ATPase-mediated biogenesis of the Golgi and ER. Symposium 'Organelle assembly and dysfunction' Invited International conference
Kondo H.
第59回日本細胞生物学会・第40回日本発生生物学会 合同大会 2007.5
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ゴルジ体・小胞体の形成と維持のための分子機構を探る Invited International conference
近藤久雄
第4回可視化技術ワークショップ 2007.11
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p97ATPaseによる細胞内膜融合とユビキチン、膜輸送をめぐるユビキチン機能の新展開 Invited International conference
近藤久雄
第30回日本分子生物学会年会・第80回日本生化学会大会 合同大会 2007.12
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p97-mediated biogenesis of the Golgi and ER Invited International conference
近藤久雄
第31回日本分子生物学会年会・第81回日本生化学会大会 合同大会 2008.12
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p97ATPaseによる小胞体・ゴルジ体の形成維持機構 Invited International conference
近藤久雄
第61回日本細胞生物学会大会 2009.6
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p97ATPase-mediated Golge and ER biogenesis
近藤 久雄
オルガネラ研究会2013 2014.2
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p97ATPase-mediated Golge biogenesis
近藤 久雄
Organella-kickoff2013 2013.3
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p97ATPase膜融合機構
近藤 久雄
第５回GCOE事業推進担当者会議 2012.2
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VCIP135 and its binding protein,WAC,in p97ATPase-membrane fusion Invited
近藤 久雄
GCOE-SU2011 2011.8
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p97ATPase-mediated membrane fusion Invited International conference
近藤 久雄
The 8th Global COE International Symposium Kyushu University - National University of Singapore Bilateral Workshop 2011.8
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試験管内再構成系を用いた小胞体形成因子の探索 Invited
近藤 久雄
第4回GCOE事業推進担当者会議 2011.2
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p97-midiated biogenesis of the Golgi and ER Invited International conference
近藤 久雄
The 20th Hot Spring Harbor Symposium joint with the 6th Global COE International Symposium 2010.8
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細胞周期における細胞内小器官ゴルジ体・小胞体の形成維持機構 Invited
近藤 久雄
第３回GCOE事業推進担当者会議 2010.2
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Cell proliferation and Death Invited International conference
近藤 久雄
GCOE 2008 2008.11
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試験管内における小胞体の網状構造の再構成 Invited
近藤 久雄
第２回GCOE事業推進担当者会議 2009.2
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細胞周期における細胞内小器官ゴルジ体・小胞体の形成維持機構 Invited
近藤 久雄
第1回GCOE事業推進担当者会議 2008.2
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p97-midiated biogenesis of the Golge and ER Invited
近藤 久雄
細胞生物学会2007 2008.10
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p97ATPase-mediated membrane fusion Invited
近藤 久雄
九大細胞生物2010 2010.9
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VCIP135 deubiquitinase and its binding protein, WAC, in p97 ATPase-mediated membrane fusion. Invited International conference
近藤 久雄, 十津川 剛
The 10th Global COE International Symposium “Biochemistry and Cell Biology” 2011.12
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p97 ATPase-mediated biogenesis of the ER and Golgi. Invited International conference
近藤 久雄, 金子 弥生
The 5th Global COE International Symposium Cell Cycle and Differentiation 2010.2
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VCIP135phosphorylation on Threonine-760 and Serine-767 is important for Golgi disassembly at mitosis International conference
近藤 久雄, 久保田章乃
Post-GCOE Symposium and Retreat in Singapore 2013.3
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p55 is a novel essential factor for p97ATPase-mediated membrane fusion in the Golgi Invited International conference
近藤 久雄, 金子 弥生
Post-GCOE Symposium in Melbourne 2014.2
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細胞内膜融合機構ｐ９７/ｐ４７経路の新規因子
近藤 久雄, 金子 弥生
第2回生命科学系GCOEネットワークフォーラム 2010.2
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p97ATPase-mediated membrane fusion Invited International conference
近藤 久雄
九大細胞生物2010 2010.9
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VCIP135phosphorylation on Threonine-760 and Serine-767 is important for Golgi disassembly at mitosis International conference
近藤 久雄, 田中愛康
Recent Advances in Stem Cell Biology 2013 2013.11
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Identification of a novel factor in p97/p47-mediated Golgi membrane fusion pathway. Invited International conference
近藤 久雄, 田村かおり
The 6th Young Investigators Forum 2011.2
MISC
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p97/p47-Mediated Biogenesis of Golgi and ER.
Uchiyama K., Kondo H.
J Biochem 2005.1
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ゴルジ体・小胞体形成に必須なp97膜融合機構
近藤久雄、内山圭司、沖米田司
細胞における蛋白質の一生（共立出版） 2004.1
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ゴルジ体の層板形成の分子機構
近藤久雄
核ー細胞質間輸送と小胞輸送（羊土社） 1999.1
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廃用性筋萎縮における酸化的ストレス
近藤久雄
活性酸素と運動（共立出版） 1999.1
Professional Memberships
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日本分子生物学会
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日本生理学会
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日本細胞生物学会
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日本生化学学会
Committee Memberships
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日本細胞生物学会 編集委員 Domestic
2019.7 - 2024.12
Academic Activities
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Screening of academic papers
Role(s)： Peer review
2024 - 2025
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Screening of academic papers
Role(s)： Peer review
2023
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Screening of academic papers
Role(s)： Peer review
2022
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Screening of academic papers
Role(s)： Peer review
2021
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Screening of academic papers
Role(s)： Peer review
2020
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Screening of academic papers
Role(s)： Peer review
2018
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Screening of academic papers
Role(s)： Peer review
2017
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細胞生物学会
2016.7 - 2023.12
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座長（Chairmanship）
日本分子細胞生物学会 （ Japan ） 2014.6 - 2013.12
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科学研究費補助金審査委員
Role(s)： Review, evaluation
日本学術振興会 2008.4 - 2009.3
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座長（Chairmanship）
日本分子生物学会 （ Japan ） 2005.12 - Present
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プログラム委員
日本細胞生物学会
Research Projects
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p97ATPase膜融合経路における新規の膜係留複合体の制御機構
Grant number：24K09439 2024.4 - 2027.3
Grants-in-Aid for Scientific Research Grant-in-Aid for Scientific Research (C)
近藤 久雄
Grant type：Scientific research funding
細胞内小器官ゴルジ体は「扁平膜積層構造」をとる。脊椎動物細胞では細胞分裂期にこのゴルジ体は小胞化する。この分裂期におけるゴルジ体の「小胞化－再構成」に特化して機能するp97/p47膜融合経路の解明は、脊椎動物細胞の細胞分裂・極性形成の仕組みの解明に直結すると考えられる。
p97/p47膜融合経路は不明なことが多いが、申請者はつい先頃、「p97/p47自身がFTCD-p97/p47-FTCD膜係留複合体を形成しゴルジ体膜が係留される」ことを発見した。本研究ではさらに、ゴルジ体上のFTCD受容体を同定しつつ、Rab蛋白質の関与、膜係留装置の形成・解離の制御機構の解明をも目指す。 -
p97ATPase/p47-mediated membrane tethering system
Grant number：21K06154 2021 - 2023
Japan Society for the Promotion of Science Grants-in-Aid for Scientific Research Grant-in-Aid for Scientific Research (C)
Kondo Hisao
Authorship：Principal investigator Grant type：Scientific research funding
VCIP135 is necessary for p97/p47-mediated Golgi membrane fusion. Although VCIP135 is known to form a complex with p97 in the cytosol, the role of this complex in Golgi and ER biogenesis has remained unclear. In this study, we demonstrated that VCIP135 has two distinct p97-binding sites at its N- and C-terminal regions. We also clarified that the N- and C-terminal binding sites in VCIP135 interact with the C- and N-terminal regions of p97, respectively. These two interactions within the complex are synchronously controlled by the nucleotide state of p97. We next generated VCIP135 mutants lacking each of the p97-binding sites to investigate their functions in living cells, and clarified that VCIP135 is involved in Golgi and ER biogenesis through its two distinct interactions with p97. VCIP135 is hence a unique p97-binding protein that functions by interacting with both the N-and C-terminal regions of p97, which strongly suggests that it plays crucial roles in p97-mediated events.
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ゴルジ体形成に働くp97ATPase膜融合のユビキチン化による制御
Grant number：16K07353 2016 - 2018
Japan Society for the Promotion of Science Grants-in-Aid for Scientific Research Grant-in-Aid for Scientific Research (C)
Authorship：Principal investigator Grant type：Scientific research funding
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細胞内小器官の異常を来す分子機構とその病態生理意義
2013 - 2014
第一三共生命科学研究振興財団
Authorship：Principal investigator Grant type：Contract research
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p97ATPase膜融合における脱ユビキチン化酵素VCIP135の作用機序
Grant number：24370082 2012 - 2014
Japan Society for the Promotion of Science Grants-in-Aid for Scientific Research Grant-in-Aid for Scientific Research (B)
Authorship：Principal investigator Grant type：Scientific research funding
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タンパク質の生産・品質管理の場としての小胞体網状構造の形成とその意義
Grant number：22020027 2010 - 2011
Grants-in-Aid for Scientific Research Grant-in-Aid for Scientific Research on Priority Areas
Authorship：Principal investigator Grant type：Scientific research funding
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細胞内小器官ゴルジ体・小胞体の各々におけるｐ９７ＡＴＰａｓｅ膜融合機構の解明
Grant number：21370091 2009 - 2011
Japan Society for the Promotion of Science Grants-in-Aid for Scientific Research Grant-in-Aid for Scientific Research (B)
Authorship：Principal investigator Grant type：Scientific research funding
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タンパク質の生産・品質管理の場としての小胞体の網状構造の意義
Grant number：20059025 2008 - 2009
Grants-in-Aid for Scientific Research Grant-in-Aid for Scientific Research on Priority Areas
Authorship：Principal investigator Grant type：Scientific research funding
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細胞内小器官ゴルジ体・小胞体の形成維持のための分子機構
Grant number：19370088 2007 - 2008
Japan Society for the Promotion of Science Grants-in-Aid for Scientific Research Grant-in-Aid for Scientific Research (B)
Authorship：Principal investigator Grant type：Scientific research funding
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新規の細胞内膜融合機構p97/p37経路の解明
Grant number：18050046 2006 - 2007
Grants-in-Aid for Scientific Research Grant-in-Aid for Scientific Research on Priority Areas
Authorship：Principal investigator Grant type：Scientific research funding
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小胞体の網状構造の形成機構
Grant number：17028060 2005 - 2006
Grants-in-Aid for Scientific Research Grant-in-Aid for Scientific Research on Priority Areas
Authorship：Principal investigator Grant type：Scientific research funding
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細胞内小器官ゴルジ体・小胞体の形態異常と病態
2002 - 2005
Grants-in-Aid for Scientific Research 科学技術振興機構・発展推進事業
Authorship：Principal investigator Grant type：Competitive funding other than Grants-in-Aid for Scientific Research
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Grant number：18K19747
金子 弥生, 近藤 久雄
Grant type：Scientific research funding
廃用性筋萎縮における酸化的ストレスの筋萎縮促進機序を明らかにすべく、細胞生物学的な観点から研究を進めた。廃用性筋萎縮における細胞内構造の変化の1つとして、ゴルジ体の小胞化を観察している。ゴルジ体の高次構造は機能のために重要だと考えられており、ゴルジ体の小胞化はその機能が損なわれていることを示唆している。ゴルジ体の小胞化には、膜融合の阻害が機序に考えられる。本研究では、膜融合の選択性に最も重要と考えられている【小胞とゴルジ体膜を繋ぎとめる係留装置】に焦点を当ててた。その成果として、①新規の膜係留因子FTCDの発見と②膜係留の観察を可能にするin vivo, in vitro両系の確立に成功した。
Class subject
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分子細胞生物Ｉ
2024.10 - 2025.3 Second semester
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統合基礎生命科学
2024.10 - 2025.3 Second semester
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生命医科学特論I
2024.4 - 2025.9 First semester
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生命医科学II
2024.4 - 2025.9 First semester
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分子細胞生物Ｉ
2023.10 - 2024.3 Second semester
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統合基礎生命科学
2023.10 - 2024.3 Second semester
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生命医科学II
2023.4 - 2023.9 First semester
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生命医科学特論I
2023.4 - 2023.9 First semester
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統合基礎生命科学
2022.10 - 2023.3 Second semester
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分子細胞生物Ｉ
2021.10 - 2022.3 Second semester
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統合基礎生命科学
2021.10 - 2022.3 Second semester
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統合基礎生命科学
2020.10 - 2021.3 Second semester
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分子細胞生物Ｉ
2020.10 - 2021.3 Second semester
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統合基礎生命科学
2019.10 - 2020.3 Second semester
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分子細胞生物Ｉ
2019.10 - 2020.3 Second semester
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細胞生物学
2019.4 - 2019.9 First semester
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分子細胞生物Ｉ
2018.10 - 2019.3 Second semester
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統合基礎生命科学
2018.10 - 2019.3 Second semester
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細胞生物学
2018.4 - 2018.9 First semester
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統合基礎生命科学
2017.10 - 2018.3 Second semester
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分子細胞生物Ｉ
2017.10 - 2018.3 Second semester
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生命医科学特講
2017.4 - 2018.3 Full year
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細胞生物学
2017.4 - 2017.9 First semester
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分子細胞生物Ｉ
2016.10 - 2017.3 Second semester
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統合基礎生命科学
2016.10 - 2017.3 Second semester
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細胞工学基礎
2016.10 - 2017.3 Second semester
-
生命医科学特講
2016.4 - 2017.3 Full year
-
細胞工学特論
2016.4 - 2016.9 First semester
-
細胞生物学
2016.4 - 2016.9 First semester
-
総合生命科学Ｉ
2015.10 - 2016.3 Second semester
-
細胞工学基礎
2015.10 - 2016.3 Second semester
-
分子細胞生物Ｉ
2015.10 - 2016.3 Second semester
-
生命医科学特講
2015.4 - 2016.3 Full year
-
細胞生物学
2015.4 - 2015.9 First semester
-
細胞工学特論
2015.4 - 2015.9 First semester
-
総合生命科学Ｉ
2014.10 - 2015.3 Second semester
-
分子細胞生物Ｉ
2014.10 - 2015.3 Second semester
-
細胞工学基礎
2014.10 - 2015.3 Second semester
-
生命医科学特講
2014.4 - 2015.3 Full year
-
細胞工学特論
2014.4 - 2014.9 First semester
-
細胞工学基礎
2013.10 - 2014.3 Second semester
-
分子細胞生物Ｉ
2013.10 - 2014.3 Second semester
-
総合生命科学Ｉ
2013.10 - 2014.3 Second semester
-
生命医科学特講
2013.4 - 2014.3 Full year
-
細胞工学特論
2013.4 - 2013.9 First semester
-
コアセミナー
2013.4 - 2013.9 First semester
-
医学入門医学生物学
2013.4 - 2013.9 First semester
-
細胞工学基礎
2012.10 - 2013.3 Second semester
-
分子細胞生物Ｉ
2012.10 - 2013.3 Second semester
-
総合生命科学Ｉ
2012.10 - 2013.3 Second semester
-
生命医科学特講
2012.4 - 2013.3 Full year
-
細胞工学特論
2012.4 - 2012.9 First semester
-
コアセミナー
2012.4 - 2012.9 First semester
-
医学入門医学生物学
2012.4 - 2012.9 First semester
-
細胞工学基礎
2011.10 - 2012.3 Second semester
-
分子細胞生物Ｉ
2011.10 - 2012.3 Second semester
-
総合生命科学セミナーI
2011.10 - 2012.3 Second semester
-
総合生命科学セミナーI
2011.10 - 2012.3 Second semester
-
細胞工学特論
2011.4 - 2011.9 First semester
-
コアセミナー
2011.4 - 2011.9 First semester
-
医学入門医学生物学
2011.4 - 2011.9 First semester
-
分子細胞生物Ｉ
2010.10 - 2011.3 Second semester
-
細胞工学基礎
2010.10 - 2011.3 Second semester
-
総合生命科学セミナーＩ
2010.10 - 2011.3 Second semester
-
国際医学
2010.10 - 2011.3 Second semester
-
細胞工学特論
2010.4 - 2010.9 First semester
-
コアセミナー
2010.4 - 2010.9 First semester
-
医学入門医学生物学
2010.4 - 2010.9 First semester
-
分子細胞生物Ｉ
2009.10 - 2010.3 Second semester
-
総合生命科学セミナーＩ
2009.10 - 2010.3 Second semester
-
細胞工学基礎
2009.10 - 2010.3 Second semester
-
細胞工学特論
2009.4 - 2009.9 First semester
-
医学入門 医学生物学
2009.4 - 2009.9 First semester
-
コアセミナー
2009.4 - 2009.9 First semester
-
細胞工学基礎
2008.10 - 2009.3 Second semester
-
総合生命科学セミナーＩ
2008.10 - 2009.3 Second semester
-
生化学
2008.4 - 2008.9 First semester
-
医学入門 医学生物学
2008.4 - 2008.9 First semester
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生命医科学特論Ⅱ
2025.6 - 2025.8 Summer quarter
-
Topics in medical life sciences Ⅱ
2025.6 - 2025.8 Summer quarter
-
生命医科学Ⅱ
2025.4 - 2025.6 Spring quarter
-
Medical Life Sciences Ⅱ
2025.4 - 2025.6 Spring quarter
-
生命医科学特論Ⅳ
2024.6 - 2024.8 Summer quarter
-
Topics in medical life sciences Ⅳ
2024.6 - 2024.8 Summer quarter
-
生命医科学Ⅱ
2024.4 - 2024.6 Spring quarter
-
Medical Life Sciences Ⅱ
2024.4 - 2024.6 Spring quarter
FD Participation
-
2024.8 Role：Participation Title：医学部FD
Organizer：[Undergraduate school/graduate school/graduate faculty]
Other educational activity and Special note
-
2010 Class Teacher 学部
-
2009 Class Teacher 学部
Outline of Social Contribution and International Cooperation activities
-
Colaboration with Imperial College group
Social Activities
-
研究指導
三菱化学生命科学研究所 2006.9
Media Coverage
-
我々が発見した細胞内構造を形成維持するための新たな仕組みの解説 Newspaper, magazine
日本工業新聞 2006.12
Travel Abroad
-
Staying countory name 1：United Kingdom Staying institution name 1：ケンブリッジ大学