Updated on 2025/06/13

写真a

 
HONJOH KEN-ICHI
 
Organization
Faculty of Agriculture Department of Bioscience and Biotechnology Associate Professor

Graduate School of Bioresource and Bioenvironmental Sciences Department of Bioscience and Biotechnology(Concurrent)
School of Agriculture Department of Bioresource and Bioenvironment(Concurrent)
Title
Associate Professor
Contact information
メールアドレス
Tel
0928024758
Profile
Some plant species develop freezing tolerance when exposed to low, non-freezing temperatures. If freezing tolerance could be provided to cold-sensitive plants, the contribution to stable production of crops would be significant. In our laboratory, we found that hardened cells of Chlorella vulgaris C-27 survive slow freezing to -196°C. We are investigating the mechanism of the development of freezing tolerance of Chlorella. Furthermore, based on the research of Chlorella, enhancement of freezing tolerance of higher plants is also carried out.
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Research Areas

  • Life Science / Food sciences

Degree

  • Ph.D of Agriculture

Research History

  • Kyushu University Faculty of Agriculture Department of Bioscience and Biotechnology  Associate Professor 

    2007.3 - Present

Education

  • Kyushu University   Graduate School, Division of Agriculture   Department of Food Science and Technology

    - 1994

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  • Kyushu University   農学研究科   食糧化学工学専攻

    - 1994

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    Country:Japan

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  • Kyushu University   農学研究科   食糧化学工学

    - 1992

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    Country:Japan

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  • Kyushu University   Graduate School, Division of Agriculture  

    - 1992

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  • Kyushu University   School of Agriculture   食糧化学工学

    - 1990

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    Country:Japan

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Research Interests・Research Keywords

  • Research theme: 食品保蔵学

    Keyword: 植物の凍結耐性、腐敗菌制御

    Research period: 2024 - Present

  • Research theme: Food Hygienic Chemistry

    Keyword: 食中毒細菌制御

    Research period: 2024 - Present

  • Research theme: Food Preservation

    Keyword: Freezing tolerance of plants, control of spoilage bacteria

    Research period: 2024

  • Research theme: Food Hygienic Chemistry

    Keyword: Food Hygienic Chemistry

    Research period: 2024

  • Research theme: Development of a method for lowering contamination with Listeria to fresh produce during cultivation

    Keyword: Listeria monocytogenes, fresh produce, decontamination

    Research period: 2013.5 - 2019.3

  • Research theme: Studies on new compatible solutes for freezing tolerance

    Keyword: compatible solutes, freezing tolerance, taurine, tolehalose

    Research period: 2009.4

  • Research theme: Studies of enhancement of freezing tolerance of Saccharomyces cerevisiae

    Keyword: freezing tolerance, Saccharomyces cerevisiae

    Research period: 1996.4

  • Research theme: Studies on the enhancement of freezing tolerance of higher plants

    Keyword: freezing tolerance, Arabidopsis thaliana, lettuce, plants

    Research period: 1994.4

  • Research theme: Studies on the mechanisms of the development of freezing tolerance of chlorella

    Keyword: chlorella, freezing tolerance, hardening

    Research period: 1991.4 - 2017.3

  • Research theme: Biocontrol of food poinsoning- and spoilage-bacteria by bacteriophages

    Keyword: バクテリオファージ、腐敗細菌、食中毒細菌

    Research period: - Present

Awards

  • 平成25年度日本食品科学工学会奨励賞

    2013.8   公益財団法人 日本食品科学工学会  

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    低温流通食品の品質と安全性確保に関する基礎的研究

Papers

  • Antibacterial and Antibiofilm Efficacy of Endolysin LysEf-9 and Crude Bacteriocin Against Multidrug and Phage-Resistant Enterococci International coauthorship

    El-Telbany, M; Wang, C; Clokie, M; Lwin, SZC; Abdelaziz, MNS; Maung, AT; Linn, KZ; Damaso, CH; Lin, YZ; Ye, HM; Masuda, Y; Honjoh, KI; Miyamoto, T

    PROBIOTICS AND ANTIMICROBIAL PROTEINS   2025.6   ISSN:1867-1306 eISSN:1867-1314

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    Language:English   Publishing type:Research paper (scientific journal)   Publisher:Probiotics and Antimicrobial Proteins  

    A gene encoding endolysin LysEf-9 from the phage PEF9, a lytic enterococcal phage, was cloned and overexpressed in Escherichia coli. The expressed LysEf-9 protein with a concentration of 62.5 μg/mL reduced the turbidity and viability of E. faecalis JCM 7783 after 3 h of incubation. LysEf-9 showed stability at different pH (2–12) and withstood different temperature treatments up to 60 °C. In addition, it showed a wide spectrum against Enterococci. Phage vB_EfKS5 with antibacterial and antibiofilm properties against E. faecalis was isolated previously. The phage-resistant mutant was isolated and used as an indicator strain in the present study. A bacteriocin-producing E. faecalis strain was also isolated, and the cell-free supernatant (crude bacteriocin) was partially purified by ammonium sulfate precipitation. The crude bacteriocin showed a wide spectrum against Enterococcus spp. and lysed the phage-resistant mutant. Using LysEf-9 with the crude bacteriocin reduced the viable count of E. faecalis JCM 3378 by 2.45 log after 24 h. In addition, LysEf-9 with phage vB_EfKS5 retarded the growth of the phage-resistant mutant. The antibiofilm efficacy of LysEf-9 and crude bacteriocin was also determined. LysEf-9 lowered the viable count of E. faecalis JCM 7783, E. faecium No. 26, and the phage-resistant mutant in the 48-h biofilm by 3 log, 2 log, and 1.5 log, respectively. The combination of LysEf-9 with crude bacteriocin reduced the total viable count of the indicator strains by at least 3 logs. This study shows the strong antibacterial and antibiofilm activity of LysEf-9, and the activity was enhanced when used with crude bacteriocin or phage.

    DOI: 10.1007/s12602-025-10616-0

    Web of Science

    Scopus

    PubMed

  • Identification of Low Temperature Inducible Genes of<i> Lactuca</i><i> sativa</i> by Using Suppression Subtractive Hybridization Method

    Honjoh, K; Okano, H; Sasaki, M; Kurokawa, M; Kimura, T; Shibata, K; Masuda, Y; Miyamoto, T

    JOURNAL OF THE FACULTY OF AGRICULTURE KYUSHU UNIVERSITY   69 ( 1 )   2024   ISSN:0023-6152

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    Language:English   Publishing type:Research paper (bulletin of university, research institution)  

    Web of Science

  • Potential application of phage vB_EfKS5 to control Enterococcus faecalis and its biofilm in food

    El-Telbany Mohamed, Lin Chen-Yu, Abdelaziz Marwa Nabil, Maung Aye Thida, El-Shibiny Ayman, Mohammadi Tahir Noor, Zayda Mahmoud, Wang Chen, Zar Chi Lwin Su, Zhao Junxin, Masuda Yoshimitsu, Honjoh Ken-ichi, Miyamoto Takahisa, El Mohamed

    AMB Express   13 ( 1 )   2023.11   eISSN:21910855

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    Language:English   Publisher:Springer  

    Contaminated food with antibiotic-resistant Enterococcus spp. could be the vehicle for transmitting Enterococcus to humans and accordingly cause a public health problem. The accumulation of biogenic amines produced by Enterococcus faecalis (E. faecalis) in food may have cytological effects. Bacteriophages (phage in short) are natural antimicrobial agents and can be used alone or in combination with other food preservatives to reduce food microbial contaminants. The aim of this study was to isolate a novel phage against E. faecalis and determine its host range to evaluate its potential application. Bacteriophage, vB_EfKS5, with a broad host range, was isolated to control the growth of E. faecalis. The vB_EfKS5 genome is 59,246 bp in length and has a GC content of 39.7%. The computational analysis of phage vB_EfKS5 genome confirmed that it does not contain any lysogenic, toxic, or virulent genes. Phage vB_EfKS5 exhibited lytic activity against most E. faecalis isolates with different multiplicities of infections and it infected 75.5% (22/29) of E. faecalis isolates and 42.3% (3/7) of E. faecium isolates. It was also able to destroy the biofilm formed by E. faecalis with different MOIs. Phage vB_EfKS5 alone or in combination with nisin could control the growth of E. faecalis in broth and milk. Based on its high productivity, stability, short latent period, and large burst size, phage vB_EfKS5 has a high potential for applications both in food and medical applications.

    CiNii Research

  • Possible Contamination Routes of Listeria monocytogenes in Leaf Lettuce during Cultivation Reviewed

    Ken-ichi Honjoh, Yin Lin, Kiyomi Jo, Yuri Iwaizako, Masayuki Maeda, Nobuyuki Kjima, Takahisa Miyamoto

    Food Sci. Technol. Res.   24 ( 5 )   911 - 920   2018.9

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    Language:English   Publishing type:Research paper (scientific journal)  

    Outbreaks of foodborne illness caused by Listeria monocytogenes in or on fresh produce have been reported. Several contamination routes have been proposed and the cultivation process is one of the suspected routes. Leaf lettuce is one of the most important fresh produce products. In the present paper, we investigated the possible routes of L. monocytogenes contamination in leaf lettuce during cultivation. Leaf lettuce was cultivated in soils inoculated with cocktails of three isolates of L. monocytogenes belonging to different serotypes (1/2a, 1/2b, and 4b). The viability and injury state of L. monocytogenes in soils, and bacteria survival in or on leaf lettuce were investigated during 10 weeks of cultivation. Soils were artificially contaminated with L. monocytogenes at levels of 4, 6 or 8 log CFU/g, followed by cultivation of leaf lettuce in the contaminated soils. Populations of L. monocytogenes in the soil decreased to less than the detection limit (< 2 log CFU/g) at 8-10 weeks after the start of cultivation. L. monocytogenes was detected in some harvested leaf lettuce leaves at low levels, almost equivalent to the detection limit. As L. monocytogenes was not detected in the leaves of leaf lettuce plants cultivated in highly-contaminated soils after surface disinfection, the possibility of internalization of L. monocytogenes into leaf lettuce plants was considered low. Spraying of water contaminated with bacteria at greater than 3.2 log CFU/plant led to the survival of bacteria on the leaf lettuce leaves even after seven days. Furthermore, leaf damage prolonged the survival period of bacteria on the leaves.

    DOI: 10.3136/fstr.24.911

    Repository Public URL: https://hdl.handle.net/2324/7183496

  • Freezing tolerance of Lactuca sativa and induction of CBF and GolS genes during cold treatment

    @Ken-ichi HONJOH, #Hitomi OKANO, #Aya KAWABATA, #Masaru KUROKAWA, #Taiki KIMURA, Takeshi MACHIDA, @Yoshimitsu MASUDA and @Takahisa MIYAMOTO

    Journal of Faculty of Agriculture, Kyushu University   63 ( 2 )   249 - 257   2018.9

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    Language:English   Publishing type:Research paper (bulletin of university, research institution)  

    Repository Public URL: https://hdl.handle.net/2324/1955390

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Books

  • Chloroplast NADPH-dependent thioredoxin reductase from Chlorella vulgaris functions as an antioxidant system with 2-Cys peroxiredoxin; The 5th International Joint Symposium between Japan and Korea. The Recent Status and Perspective of Food System, Agricultural Environment and Biology

    Takeshi Machida, Akiko Ishibashi, Eri Kato, Jun-ichi Sato, Shinji Kawasaki, Youichi Niimura, Ken-ichi Honjoh and Takahisa Miyamoto(Role:Joint author)

    2008.12 

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    Language:English   Book type:Scholarly book

  • 植物の耐凍性向上への試み、生物機能研究の進歩1,

    本城賢一(Role:Joint author)

    生物機能研究会編集  2002.3 

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    Language:Japanese   Book type:Scholarly book

  • 食糧の安定供給をめざして:植物への耐凍性付与、生物機能の新展開-人類の生存をかけて-

    本城賢一(Role:Joint author)

    日本学術会議 栄養・食糧科学研究連絡委員会編  2000.11 

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    Language:Japanese   Book type:Scholarly book

  • 食品製造・検査における芽胞・損傷菌とその検出・制御技術

    木嶋伸行、本城賢一、前田征之(Role:Joint author)

    シーエムシー出版  2020.5 

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    Responsible for pages:315-318   Language:Japanese   Book type:Scholarly book

Presentations

  • レタス由来 LEA タンパク質発現大腸菌のストレス耐性

    鍵谷俊太、新名祐作、本城賢一、益田時光、宮本敬久

    2025年度日本栄養・食糧学会九州・沖縄支部 日本食品科学工学会 西日本支部 合同大会  2024.11  日本栄養・食糧学会九州・沖縄支部 日本食品科学工学会 西日本支部

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    Event date: 2024.11

    Language:Japanese   Presentation type:Oral presentation (general)  

    Venue:長崎   Country:Japan  

  • 低温貯蔵と不十分な加熱がカンピロバクターの生残性に及ぼす影響

    北山玲衣、本城賢一、益田時光、宮本敬久

    2025年度日本栄養・食糧学会九州・沖縄支部 日本食品科学工学会 西日本支部 合同大会  2024.11  日本栄養・食糧学会九州・沖縄支部 日本食品科学工学会 西日本支部

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    Event date: 2024.11

    Language:Japanese   Presentation type:Oral presentation (general)  

    Venue:長崎   Country:Japan  

  • Treating multidrug and phage-resistant Enterococcus spp. by phage cocktails, endolysin, and bacteriocin International coauthorship International conference

    Mohamed El-Telbany, Chen Wang, Ayman El-Shibiny, Yoshimitsu Masuda , Ken-ichi Honjoh , Takahisa Miyamoto

    Viruses of Microbes 2024  2024.7  International Society for Viruses of Microorganisms

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    Event date: 2024.7

    Language:English   Presentation type:Poster presentation  

    Venue:Cairns   Country:Australia  

  • 抗菌剤散布がレタス栽培土壌中の薬剤耐性菌に及ぼす影響

    安留正倫、中尾朱李、實政裕哉、益田時光、本城賢一、 木嶋伸行、石井孝昭、宮本敬久

    第61回化学関連支部合同九州大会  2024.6  日本農芸化学会西日本支部その他

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    Event date: 2024.6

    Language:Japanese   Presentation type:Poster presentation  

    Venue:北九州市   Country:Japan  

  • 空芯菜由来抗菌タンパク質の探索

    ○實政裕哉、北山玲衣、中川義健、益田時光、本城賢一、宮本敬久

    令和5年度 日本食品科学工学会 西日本支部大会  2023.10 

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    Event date: 2023.10

    Language:Japanese   Presentation type:Oral presentation (general)  

    Venue:福岡市(九州大学西新プラザ)   Country:Japan  

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MISC

  • 糖修飾LEAタンパク質の利用を基盤とする食肉の新しい凍害防御技術の確立

    本城賢一, 井倉則之, 野間誠司

    2012.11

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    Language:Japanese  

  • 生鮮野菜の凍結貯蔵を意図した分子生物学的基礎研究

    本城賢一

    2012.10

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    Language:Japanese  

  • 九大農学部の安全の取り組み

    本城賢一

    日本安全学教育研究会誌   2011.8

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    Language:Japanese   Publishing type:Internal/External technical report, pre-print, etc.  

  • 植物の耐凍性獲得に関わる新規抗酸化系酵素群の解析

    本城賢一

    2009.5

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    Language:Japanese  

  • トレハロース蓄積による酵母の耐凍性向上の試み

    本城賢一

    2009.2

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    Language:Japanese  

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Professional Memberships

  • Japanese Society for Food Hygiene and Safety

    2023.12 - Present

  • Japanese Society for Safety Education

    - Present

  • The Japanese Society of Plant Physiologists

    - Present

  • The Japanese Society for Food Science and Technology

    - Present

  • Japan Society for Bioscience, Biotechnology, and Agrochemistry

    - Present

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Committee Memberships

  • 日本食品衛生学会   学会活性化委員会委員   Domestic

    2025.4 - Present   

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    Committee type:Academic society

  • 日本食品科学工学会   国際交流委員会副委員長   Domestic

    2024.3 - Present   

  • 日本食品科学工学会   西日本支部代議員   Domestic

    2022.3 - Present   

  • 日本農芸化学会   広報委員   Domestic

    2021.3 - 2025.2   

  • 日本食品科学工学会   総務委員会委員   Domestic

    2020.4 - Present   

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Academic Activities

  • 日本食品科学工学会 第68回大会(オンライン) 庶務 International contribution

    Role(s): Planning, management, etc.

    ( Japan ) 2021.8

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    Type:Competition, symposium, etc. 

    Number of participants:1,000

  • 日本食品科学工学会 第67回大会 庶務

    (コロナウイルス感染拡大のため中止)  ( Japan ) 2020.8

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    Type:Competition, symposium, etc. 

    Number of participants:0

  • 日本食品科学工学会第61回大会 庶務

    Role(s): Planning, management, etc.

    ( Japan ) 2014.8

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    Type:Competition, symposium, etc. 

    Number of participants:1,100

  • 第33回日本食品微生物学会学術総会 庶務

    Role(s): Planning, management, etc., Panel moderator, session chair, etc.

    ( Japan ) 2012.10

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    Type:Competition, symposium, etc. 

    Number of participants:600

  • 日本農芸化学会1999年大会 庶務 International contribution

    Role(s): Planning, management, etc.

    日本農芸化学会1999年度大会  ( Japan ) 1999.3

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    Type:Competition, symposium, etc. 

    Number of participants:5,000

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Research Projects

  • バンコマイシン耐性腸球菌に作用する新規ファージの単離とならびにその特性評価によるバイオコントロール法の確立

    2025.5 - 2026.3

    公益財団法人江頭ホスピタリティ事業振興財団 令和7年度 研究開発助成

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    Authorship:Principal investigator  Grant type:Competitive funding other than Grants-in-Aid for Scientific Research

  • 環境への抗菌剤・薬剤耐性菌の拡散量低減を目指したワンヘルス 推進プロジェクト 小課題4 農薬としての抗菌剤のほ場投入による土壌及び野菜由来微生物の薬剤 耐性化機構の解明 実行課題 410:土壌環境における農薬残布の耐性菌出現への影響解明

    Grant number:JPJ008617. 22682153   2025.4 - 2026.3

    農林水産省  Research commissions

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    Authorship:Coinvestigator(s)  Grant type:Other funds from industry-academia collaboration

  • 寄附金

    2025

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    Grant type:Donation

  • 環境への抗菌剤・薬剤耐性菌の拡散量低減を目指したワンヘルス 推進プロジェクト 小課題4 農薬としての抗菌剤のほ場投入による土壌及び野菜由来微生物の薬剤 耐性化機構の解明 実行課題 410:土壌環境における農薬残布の耐性菌出現への影響解明

    Grant number:JPJ008617. 22682153  2024.4 - 2025.3

    農林水産省  Research commissions

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    Authorship:Coinvestigator(s)  Grant type:Other funds from industry-academia collaboration

  • Functional analysis of cold-inducible VPE gene and development of freeze-resistant lettuce based on the analysis

    Grant number:24K08764  2024 - 2026

    Japan Society for the Promotion of Science  Grants-in-Aid for Scientific Research  Grant-in-Aid for Scientific Research (C)

    本城 賢一, 後藤 栄治, 益田 時光

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    Authorship:Principal investigator  Grant type:Scientific research funding

    これまでに申請者らは、生鮮野菜であるレタスが栽培段階で3℃の低温処理を施されることにより、-2から-3℃での凍結・解凍にも耐える“凍結耐性”を獲得することを明らかにした。その際、プログラム細胞死に関わり凍結耐性に「マイナス」に働くと推定される液胞プロセッシング酵素(VPE)遺伝子が低温処理時に高発現し、レタスの凍結耐性レベルに影響している可能性を見出した。そこで本研究では、VPE欠損シロイヌナズナを用いてVPEと凍結耐性との関係を明らかにし、さらに、VPE遺伝子をノックアウトしたレタスの作出により凍結貯蔵性を高めたより高品質な生鮮野菜の提供に資する研究を実施する。

    CiNii Research

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Educational Activities

  • 学部学生に対しては「食品衛生化学」、「食品保蔵学」、「食品加工学」の講義を行い,「分析化学実験(分担)」,「食品衛生化学実験」の実習指導を行っている。また、国際コースにおいては「Bio-engineering(分担)」を担当している。
    大学院の修士課程学生に対しては「食品加工学特論(分担)」、「食品品質学特論(EJ科目、分担)」の講義を担当している。国際コースにおいては「Food Science and Food Technology(分担)」を担当している。

Class subject

  • 食品加工学

    2025.12 - 2026.2   Winter quarter

  • 食品衛生化学実験

    2025.10 - 2026.3   Second semester

  • 分析化学実験(食糧化学工学分野)

    2025.10 - 2026.3   Second semester

  • Food Science and Food Technology (分担)

    2025.10 - 2025.12   Fall quarter

  • 食品保蔵学

    2025.10 - 2025.12   Fall quarter

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FD Participation

  • 2021.7   Role:Participation   Title:農学研究院FD「科研費を獲りにいこう! 科研費獲得の技術と工夫」

    Organizer:[Undergraduate school/graduate school/graduate faculty]

  • 2019.5   Role:Participation   Title:優良な博士人材の獲得と育成に向けて~農学研究院教授に学ぶ ~

    Organizer:[Undergraduate school/graduate school/graduate faculty]

  • 2017.9   Role:Participation   Title:研究分析ツール「SciVal」及び研究者プロファイリングツール「Pure」に関する説明会

    Organizer:[Undergraduate school/graduate school/graduate faculty]

  • 2017.7   Role:Participation   Title:「奨学金プログラムについて」

    Organizer:[Undergraduate school/graduate school/graduate faculty]

  • 2015.9   Role:Participation   Title:「ハラスメントの実際と対応」ハラスメントの事例を通じた相談や苦情申立の流れについて

    Organizer:[Undergraduate school/graduate school/graduate faculty]

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Visiting, concurrent, or part-time lecturers at other universities, institutions, etc.

  • 2020  中村学園大学大学院栄養科学部  Classification:Part-time lecturer  Domestic/International Classification:Japan 

    Semester, Day Time or Duration:前期、水曜日、5時限

  • 2016  放送大学  Classification:Intensive course  Domestic/International Classification:Japan 

    Semester, Day Time or Duration:2016.12.17-18

  • 2009  福岡女子大学  Classification:Part-time lecturer  Domestic/International Classification:Japan 

    Semester, Day Time or Duration:火曜日4限,前期

  • 2008  福岡女子大学  Classification:Part-time lecturer  Domestic/International Classification:Japan 

    Semester, Day Time or Duration:木曜日3限,後期

  • 2007  福岡女子大学  Classification:Part-time lecturer  Domestic/International Classification:Japan 

    Semester, Day Time or Duration:木曜日3限,後期

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Participation in international educational events, etc.

  • 2018.1

    九州大学

    国際インバウンドプログラム (実験担当)

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    Number of participants:10

  • 2017.1

    九州大学

    国際インバウンドプログラム (実験担当)

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    Number of participants:16

Other educational activity and Special note

  • 2025  Coaching of Students' Association  学友会剣道部

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    部長

  • 2024  Coaching of Students' Association  学友会剣道部

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    部長

  • 2024  Development of Textbook and Teaching Material  食品衛生学 第3版 (東京化学同人)

  • 2023  Coaching of Students' Association  学友会剣道部

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    部長

  • 2022  Class Teacher  学部

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Social Activities

  • 第144回農芸化学サイエンスカフェin福岡

    日本農芸化学会  福岡市科学館  2022.12

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    Audience:General, Scientific, Company, Civic organization, Governmental agency

    Type:Lecture

  • 第138回農芸化学サイエンスカフェin福岡

    日本農芸化学会  福岡市科学館  2021.12

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    Audience:General, Scientific, Company, Civic organization, Governmental agency

    Type:Lecture

  • 講演タイトル「野菜生産段階におけるリステリアの汚染機構について」

    食品の品質保証懇話会 九州部会定例会  オンライン  2021.8

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    Audience:General, Scientific, Company, Civic organization, Governmental agency

    Type:Lecture

  • 食品の品質保持ならびに安全性確保に向けた生物学的知識の活用/システム生命科学夏の学校2019

    九州大学大学院システム生命科学府  九州大学伊都キャンパスウエスト5号館226室  2019.7

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    Audience:General, Scientific, Company, Civic organization, Governmental agency

    Type:Lecture

  • 食品の品質保持ならびに安全性確保に向けた生物学的知識の活用/システム生命科学夏の学校2019

    九州大学大学院システム生命科学府  九州大学伊都キャンパスウエスト5号館226室  2019.7

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    Type:Lecture

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Media Coverage

  • 農作物の凍結保存技術の開発について Newspaper, magazine

    2013.9

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    農作物の凍結保存技術の開発について

Activities contributing to policy formation, academic promotion, etc.

  • 2024.7 - Present   福岡県

    福岡県食品安全・安心委員会委員

  • 2020.7 - Present   福岡市

    福岡市食の安全安心推進協議会委員

Acceptance of Foreign Researchers, etc.

  • University of Sadat City

    Acceptance period: 2022.5 - 2022.10   (Period):1 month or more

    Nationality:Egypt

    Business entity:Foreign governments, foreign research institutes, international organizations

  • カントー大学

    Acceptance period: 2019.7 - 2019.8   (Period):1 month or more

    Nationality:Viet Nam

    Business entity:Foreign governments, foreign research institutes, international organizations

  • Vietnam National University of Agriculture

    Acceptance period: 2016.4 - 2016.9   (Period):1 month or more

    Nationality:Viet Nam

  • 瀋陽師範大学

    Acceptance period: 2013.7 - 2013.9   (Period):1 month or more

    Nationality:China

    Business entity:Government agency

  • カセサート大学

    Acceptance period: 2012.2 - 2012.8   (Period):1 month or more

    Nationality:Thailand

    Business entity:Foreign governments, foreign research institutes, international organizations

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Travel Abroad

  • 2013.9

    Staying countory name 1:Singapore   Staying institution name 1:MAX Atria, Singapore Expo

  • 2007.1

    Staying countory name 1:Germany   Staying institution name 1:オスナブリュック大学

  • 2000.2 - 2001.2

    Staying countory name 1:United States   Staying institution name 1:アリゾナ大学